Archive for Round Oak Rag Apple Elevation

24 Cows Built Half the Holstein Breed. The Other Half Came from One Ontario Barn.

Twenty-four cows nobody else wanted from Kansas. One stubborn Ontario dairyman who used an outcross bull exactly once. Between them: 99.84% of every A.I. sire in your barn tonight.

In 1947, at the Wisconsin State Reformatory in Green Bay, a bull calf was loaded onto a truck bound for Kansas, and not one dollar changed hands.

The Reformatory ran the best Holstein herd of all the Wisconsin state institutions — and yes, that’s exactly what it sounds like. A prison. With a bull barn that half the Midwest wanted access to. Their blood went into serious herds, the kind of operations that produced bulls like Osborndale Ivanhoe, and a breeder who got access to it got it because somebody there thought well of him.

Elmer Dawdy had no money to speak of.

What he had was a young bull sired by Dunloggin Golden Cross — and he’d gotten that mating almost by accident. Golden Cross spent fourteen days at Dawdy’s place while en route to Colorado, and Dawdy bred one cow to him during the stopover. One cow. That was the whole opportunity, and he took it.

So he offered that son to the Reformatory, straight across, for a five-week-old calf named Wis Burke Ideal, born May 20, 1947.

They took the deal.

A five-week-old calf. Understand what that means — Dawdy wasn’t buying proven genetics; he was buying a guess. Nobody knew what that calf would become. He could have grown into nothing at all and cost Dawdy the only good bull he’d ever bred by accident.

Instead, Wis Burke Ideal sired Tidy Burke Elevation. Tidy Burke Elevation sired Round Oak Rag Apple Elevation. And Elevation — the bull Holstein International crowned Bull of the Century in 1999 — sits behind roughly half of every active Holstein A.I. sire in North America today.

The other half traces to a man in Brampton, Ontario, whose Glenvue prefix Dawdy had probably never bothered to learn, whose favorite bloodline Dawdy would have thrown him out of the barn for mentioning, and who was building the identical monument from the opposite end.

ACT I: BREEDING BLIND

Understand what these men were working with, because it changes how you read everything that follows.

When Doug Dunton bought his first Holstein in 1920, formal classification didn’t exist in Canada yet. Holstein Canada wouldn’t launch its program until 1925. And even once inspectors were coming through with clipboards and scorecards, what they left behind was a letter on a card — nothing more.

No sire summaries. No daughter averages published in a booklet you could carry to a sale. No reliability figures, no indexes, no way to compare one bull’s daughters against another’s except by driving to see them.

So that’s what you did. If you wanted to know whether a bull’s daughters had udders worth having, you got in a car — or a wagon — and you went and looked. You asked the neighbors. You trusted your eye.

Doug Dunton put it plainly years later: if you couldn’t see an animal’s ancestors to judge how good the type was, and whether it passed along, “you were working blind.”

Compare that to what’s sitting in your pocket. Genomic proofs. Reliability percentages. Haplotype screens on animals that haven’t been born yet. These two men laid the foundation of the modern Holstein cow with less usable information than you’d bring to buying a used truck.

The one cow that ended the Shorthorn experiment

Doug Dunton came to Holsteins sideways, and late.

In 1920 he signed up for a dairy short course run by the Ontario Department of Agriculture. Up to then he and his father had been crossing Dual Purpose Shorthorns — an experiment the record describes, with what I read as considerable mercy, as having “little apparent success.”

During the course, Doug got into some good Holstein barns for the first time. He came home and bought one cow. One. Not a conversion — a test.

She milked 80 pounds in a day.

Twice what any Shorthorn on the place had ever given. Picture her standing in that tie-stall row, a big black-and-white stranger among the roans, filling a pail while the home-bred cows on either side of her filled half of one.

You don’t need a scorecard to read that. Doug and his father went out and bought four more Holsteins, and the Shorthorns were finished.

Three of the four came from Lorne Davidson’s farm at Meadowvale. Two of those three built everything.

Bruach Noelle hit the ground on Christmas Day, 1928 — a daughter of King Segis Vale Spofford. She’d become the maternal great-granddam of two All-Canadian winners.

Her maternal sister, Bruach Luella Acme, was a daughter of King Toitilla Acme, the Davidson herd sire. She started the line that ran down to Glenvue Jean Inka B.

Hold onto Jean Inka B. She comes back at the end, and when she does, it’s going to land.

Dunton had no college degree. What he had instead was a conviction so fixed it worked like faith. He believed in close breeding — “breeding warm,” he called it, which is honestly the most farmer description of inbreeding ever committed to print — and he believed the King Toitilla Acme line was better than whatever anyone else was using.

In roughly four decades of breeding, Doug Dunton used an outcross bull exactly once.

The verdict on that experiment is the driest sentence in the whole record: he “didn’t do much harm,” but “neither did he do much good.”

No more outcross bulls were used. Ever.

The Kansas boy who sold his herd to buy an education

Elmer Alfred Dawdy was born in 1915 on his father’s place in Washington County, north-central Kansas. He came to dairy cattle through 4-H — with a Guernsey calf, which ought to make every Holstein person reading this smile. By the time college came around, he’d built a small herd of his own.

Then he sold it. All of it. For tuition.

He sold the herd. That’s Elmer Dawdy. He’d spend his whole life treating cattle as instruments rather than ornaments, and he opened by liquidating his own to buy an animal husbandry degree from Kansas State.

He graduated in 1938 into extension work — assistant county ag rep for Saline County, then head county agent six months later. That got him named Southwestern fieldman for the Holstein Association, seven states: Texas, New Mexico, Oklahoma, Nebraska, Colorado, Wyoming, Kansas.

And at that point Elmer Dawdy had never owned a Holstein cow in his life.

Think about the strangeness of that. The Association’s man across a seven-state territory, and he didn’t own one of the animals he was paid to promote.

But the job gave him something no amount of money could buy — he now knew where every bargain, every overlooked cow, and every seller in trouble was located across a quarter of the country.

So he bought a hundred acres in Saline County and started collecting.

Crooked cows from institutions

The Tidy Burke herd never exceeded twenty-four head of milking age.

Twenty-four. Smaller than most modern hospital pens. Smaller than the string a lot of you would cull in a bad year. That’s the operation we’re talking about.

And the cows in it were, to put it charitably, crooked. Bought for a song, many carrying some physical defect — the kind of fault that makes a breeder’s eye slide right past an animal at a sale. Dawdy fixed what he could and lived with the rest.

Two came out of Clyde Hill Farm in Clyde, Missouri. One of them, Clyde Hill Jewel Carine Mercedes, was classified Good the day he bought her.

She was never raised. Not once, the entire time he owned her.

Good. In an era when Good meant middling — the score you didn’t mention at the feed store, the cow you apologized for. Year after year the classifier came through, ran his eye over her, and left the card exactly where it was.

That cow founded the family that produced Tidy Burke Elevation.

Sit with that if you’ve ever culled a heifer on her score alone. The maternal foundation behind half the breed’s paternal lineage was a permanently Good cow from a Missouri farm, bought cheap by a man who couldn’t afford better.

The rest of the foundation reads like a list of places nobody shops for cattle.

The other Clyde Hill cow became the dam of Raven Burke Elsie — three Gold Medal sons. And at the Colorado State Home for Dependent Children, Dawdy found Posch Ormsby Johanna Bess. She’d produce Ormsby Burke Pontiac Mose, sire of Paclamar Astronaut.

An orphanage. A children’s home. A prison herd. A Missouri dispersal.

Even his first herd sire came through persuasion. Dawdy talked Sister Xavier of St. Joseph’s Orphanage into buying Sir Bess Tidy from Fred Eiten of Waterloo, Iowa — Eiten proved the bull, the orphanage owned him, and Dawdy used him artificially.

A nun financed the first sire in a program that would reshape the global Holstein population, and there’s nothing in the record suggesting Dawdy ever found that arrangement remarkable.

They would not have gotten along

Doug Dunton was pleasant. Accommodating. The kind of man neighbors dropped in on without calling first. He worked as an official classifier for the Canadian association — not a job they hand to men who can’t be diplomatic in somebody else’s barn. And when his sons Ralph and Earl came of age, they became his partners.

Elmer Dawdy could be abrasive and abrupt, inclined to speak his mind regardless of consequences. His own biographer noted that his normal ways of doing business were at times “very much at odds with accepted methods and behaviours.”

His two sons left home after high school. The record’s phrasing is nine words long and quietly devastating: they couldn’t, it seemed, get away fast enough.

There’s a whole story in those nine words, and I’m not going to pretend I know it. But anyone who’s worked a farm alongside family understands what a kitchen can feel like when the man running things is certain about everything.

Then there’s the bloodline schism, which was absolute. Dunton was a King Toitilla Acme man who braided in the Rag Apple line and spent decades weaving the two together.

Dawdy was a Burke man. Tried and true.

Mention “Rag Apple” in Elmer Dawdy’s barn, and he’d throw you out.

Not argue. Throw you out.

ACT II: THE TRADE AND THE TWELVE-LACTATION COW

A row of matched half-sisters

By the mid-1950s, Glenvue Farm at Brampton was producing cows people drove to see.

There’s a photograph from those years that captures it better than any production string could — a row of A.B.C. Midnight Joe daughters standing in Dunton’s stable, and the record notes they “made an impressive sight.”

Picture what that actually means. Half-sisters, tied side by side, matched enough in their toplines and udder attachments that a visitor walking the alley could tell before anyone said a word which bull had sired the lot of them.

That’s what breeders came for. Not one great cow — anybody gets lucky once. A row of them, predictable enough to look like they’d been stamped.

The individuals were extraordinary too.

A.B.C. Inka May (EX), All-Canadian 4-year-old in 1947, made 24,142 pounds of milk at 4.67% with 1,128 pounds of fat over 365 days, milked three times a day — that’s what “3X” means on an old record, and it was the going method for pushing a cow to her limit before anyone had heard of a robot.

In 1947. When the good cow in your neighbor’s barn made 12,000, and he talked about her all winter.

Glenvue Noelle Inka (EX-4*), All-American and All-Canadian aged cow, 1947. Haltered and posed against the gate — this is what Dunton’s type-first breeding looked like when it worked. Best in her class on both sides of the border in one season, and her line ran forward to Walkway Chief Mark.

Glenvue Noelle Inka (EX-4*) took All-American and All-Canadian aged cow in 1947 — both sides of the border in one season. Glenvue Rilla Top Grade (EX) was Reserve All-Canadian aged cow in 1951.

And Glenvue Rosalie Fond Hope (VG), out of Rilla Top Grade by Spring Farm Fond Hope, stood grand champion at the Royal Winter Fair in 1955 and took All-Canadian 4-year-old the same year — on the tanbark at Toronto, in front of a crowd that considered the Royal the only show in Canada that truly counted.

Nine Glenvue matrons made Dave Morrow’s list when he asked “Who Breeds the Brood Cows?” in the June 10, 1962 Holstein World. Nine, from one farm.

And standing above all of them, an old cow named Glenvue Nettie Jemima (EX-13*).

Glenvue Nettie Jemima (EX-13*), All-Canadian aged cow, 1954. That’s not a young cow — twelve lactations wore that frame down. In a barn culling on Good Plus, Dunton chose her twelve separate times.

Twelve lactations. 218,215 pounds of milk. 8,193 pounds of fat.

All-Canadian aged cow, 1954 — in a herd where a cow graded Good Plus or she went down the road.

The number that matters there isn’t the milk. It’s the twelve.

Because Dunton ran the hardest cull rule in the country: to stay at Glenvue, a cow had to grade Good Plus or better. One exception, only one — a two-year-old who classified Good but carried a Good Plus udder got another year, because Dunton had seen too many good udders show up late.

Everything else went down the road. No sentiment, no second thoughts.

So twelve lactations in that herd means Doug Dunton stood in front of that cow twelve separate times, in an operation where merely good wasn’t good enough, and chose her. Through her prime and well past it.

A pleasant, accommodating man who culled without mercy kept looking at Nettie Jemima and deciding, again, that she stayed.

Picture her in her later years — a big, well-worn old cow with the hips and the hollows a twelve-lactation record leaves behind, still carrying an Excellent card, standing in a barn full of animals young enough to be her great-granddaughters. Anyone who’s kept an old favorite past the point where the spreadsheet says to knows exactly what that looks like.

She paid him for every one of those decisions. She’s the dam of Rosafe Centurion, and of Magician, Signet, and Citation R.

That “13*” on her card, for anyone under fifty: the star system marked qualifying superior offspring. Thirteen of them, in an era of hand-kept records and postal reporting, was a maternal record that bordered on absurd.

The cruel part of Doug Dunton’s career

Here’s what I find genuinely hard about his story.

Those cows — and type-wise they were far beyond anything Dawdy ever bred, a man who never once placed a cow on the All-American calendar — made other men famous.

A.B.C. Inka May was sold in dam — sold while carrying the calf. A.B.C. Farms bought her mother, Temple Farm May, and under the rules of the day the buyer registered the resulting offspring under his own prefix.

So the cow Doug Dunton bred carried somebody else’s name. And her son by Montvic Rag Apple Sovereign — A.B.C. Reflection Sovereign, called by some the greatest bull the breed has produced — carried it too.

Dunton bred the brood cows. Other men banked the bulls.

In his entire career, exactly one bull carrying the Glenvue prefix achieved major impact. His name was Glenvue Clipper, and he’ll matter shortly.

What he was actually selecting for

Before the trade, sit with Dunton’s philosophy a minute, because it’s the most useful thing in this story for anybody building a mating list in 2026.

“It takes longer to breed type than production,” he told a reporter, “but once you get type, it is not so difficult to get higher production.”

Type first. Always. Forced to choose, type won.

And he selected for three things that appeared nowhere on a pedigree in his day — no index, no proof, no number attached to any of them: temperament. How a cow milks out. Longevity.

Read that list again and tell me it isn’t the conversation this industry is finally having, seventy years late, about functional traits, productive life, and whether we bred ourselves into a corner chasing yield.

Dunton was selecting for traits that wouldn’t get formal evaluations for another half-century. He did it because he’d decided they mattered more than the things he could measure.

His four stated reasons for Glenvue’s consistency, from a 1963 Farmers Advocate interview: strict culling. Slow-maturing animals — his two-year-olds were never pushed, and a thousand pounds of milk in a month satisfied him. Line breeding. The Good Plus rule.

Never pushed. In a business that has spent eighty years pushing.

1947, and the fortnight bull

Down in Kansas, Dawdy had been watching Wisconsin Admiral Burke Lad tear through the breed in the early 1940s — “witnessing the havoc,” the record says — and he jumped in with both feet.

He bought Weber Burke Hazelwooder, a Burke Lad son. Then he borrowed Weber Hazelwood Burke Raven from Jake Zarnowski.

Borrowed. Neighbor to neighbor, on a handshake, the way things actually got done before contracts.

Raven was out of an inbred Burke Lad daughter, and he sired Raven Burke Elsie plus Raven Burke Carine Mercedes — the maternal granddam of Tidy Burke Elevation.

Then came the trade.

Wis Burke Ideal, born May 20, 1947, was sired by Wis Ideal (EX-GM) out of Wis Juanita — and both of them were sired by Weber Burke Cyclone, the great inbred Burke son standing at Green Bay.

A son of Cyclone bred to a daughter of Cyclone. That’s the calf Dawdy got for a bull out of a fourteen-day stopover.

Wis Burke Ideal (VG-88 GM), born May 20, 1947. Cyclone on both sides — a son of Weber Burke Cyclone bred to a Cyclone daughter. Elmer Dawdy got him from the Green Bay Reformatory as a five-week-old calf, for a bull he’d bred almost by accident. Look at the depth through that chest. Half the Holstein breed’s paternal line runs through it.

And then he doubled down. Dawdy bred Wis Burke Ideal to a daughter of Wisconsin Admiral Burke Lad — inbreeding stacked on inbreeding, on purpose, by a man whose entire philosophy on the subject ran five words long:

“You can never get too much of a good thing.”

He meant it literally. He was spectacularly right, and — as the breed would learn expensively, decades later — he was also wrong.

The calf was Tidy Burke Elevation.

August 30, 1965

On Round Oak Farm in Virginia, out of a cow named Round Oak Ivanhoe Eve by Osborndale Ivanhoe, Tidy Burke Elevation’s son hit the ground.

Round Oak Rag Apple Elevation.

Round Oak Rag Apple Elevation (EX-96 GM), b. August 30, 1965 — Bull of the Century, 10,000 A.I. sons, and 51.06% of North America’s active Holstein sire lines. His sire came out of a 24-cow Kansas herd. Read more: Round Oak Rag Apple Elevation: The Bull That Changed Everything)

And here’s the collision that makes these two men inseparable rather than rivals.

Elevation’s sire came from Dawdy. But run up the maternal line: Elevation’s maternal granddam was Round Oak Nettie Emaline, born January 7, 1959, sired by Glenafton Gaiety — and Gaiety was sired by Rosafe Centurion, the son of Dunton’s Inka Supreme Reflection out of Glenvue Nettie Jemima.

The old twelve-lactation cow. The one Doug Dunton kept choosing.

The bull named for the Rag Apple line that Elmer Dawdy would have thrown a man out of his barn for mentioning carried Doug Dunton’s brood cow on his dam’s side.

Neither man had any say in it. Neither, so far as anyone can tell, ever met the other.

May 9, 1962 — and three years earlier, Chief

May 9, 1962 — and three years earlier, Chief

Three years before Elevation, the other half of the breed was born — and not where you’d expect. Lester Fishler’s Pawnee Farm sat on the south edge of Central City, Nebraska, and when he dispersed the herd on April 14, 1962, a pregnant cow named Pawnee Farm Glenvue Beauty sold for $4,300 to Wally Lindskoog of Arlinda Farms. She rode a train to Turlock, California, and calved twenty-five days later.

The dam was Pawnee Farm Glenvue Beauty (EX) — sired by Glenvue Clipper. Dunton’s one great bull.

The sire was Pawnee Farm Reflection Admiral (born February 16, 1960, EX, Gold Medal Sire). Admiral’s sire was Rosafe Pearl Hannibal — bred on Hector Astengo’s Rosafe farm in Ontario by mating Rosafe Shamrock Rosamond to Rosafe Sovereign Supreme, which made Hannibal a double grandson of A.B.C. Reflection Sovereign.

And Rosafe Shamrock Rosamond’s maternal granddam?

Glenvue Jean Inka B.

The cow descending from Bruach Luella Acme — one of the five Holsteins a young Doug Dunton bought in 1920, after a single cow gave him 80 pounds of milk and ended his father’s Shorthorn experiment.

Pawnee Farm Arlinda Chief (EX-94), b. May 9, 1962. Sixteen thousand daughters. Five hundred thousand granddaughters. Two million great-granddaughters — and a lethal recessive nobody would find for another fifty years. Doug Dunton bred into both sides of that pedigree and never owned a share of him.

Pawnee Farm Arlinda Chief. Dunton’s breeding on both sides of the pedigree. Sire’s side and dam’s side. (Read more: The $4,300 Gamble That Reshaped Global Dairy Industry: The Pawnee Farm Arlinda Chief Story)

Elmer Dawdy contributed nothing to Chief — not one gene he was responsible for placing there.

Two bulls, born three years apart. Elevation, built on Dawdy’s paternal line with Dunton’s Nettie Jemima standing in the maternal background. Chief, built on Dunton at both ends.

That’s the climax of this story, and it took two stubborn men roughly forty-five years of refusing to compromise to arrive at it.

Two men, two halves — the ledger

 The Dawdy Side (paternal power)The Dunton Side (brood cow power)
Foundation animalWis Burke Ideal (b. May 20, 1947)Glenvue Nettie Jemima (EX-13*)
Scale / signatureNever above 24 head milkingNine great matrons on one 1962 list
Key anchor bullRound Oak Rag Apple Elevation (1965)Pawnee Farm Arlinda Chief (1962)
Paternal share, 2010s bulls51.06% of active lines48.78% of active lines
Breed contribution10,000+ A.I. sons; Bull of the Century, 1999~14% of the modern U.S. Holstein genome
The bill that came dueInbreeding velocity — “never too much of a good thing”Carried HH1, the APAF1 lethal recessive

Paternal-share figures from the Journal of Dairy Science Y-chromosome study of 62,897 North American bulls born 1950–2013. Note the irony in the bottom row: each man’s signature strength produced the other’s kind of problem.

ACT III: WHAT THEY LEFT

Neither man took a victory lap

This is the part that gets me.

In the late 1960s — right as Elevation was beginning to make his name — Elmer Dawdy got rid of his cattle. And he didn’t hold a dispersal.

Brookview Tony Charity. A half-interest in this one cow brought $1,450,000 at the 1985 Hanover Hill dispersal — 2,500 people from around the world, Bob Shore working the block, buyers leaning on the rail from three continents. This is the send-off Elmer Dawdy could have had. He handed his twenty-four cows to two young men he trusted and drove home. (Read more: Brookview Tony Charity: A $47,000 Gamble That Outlived Everyone Who Doubted Her)

Understand what that means. On July 15th and 16th, 1985, the Hanover Hill dispersal at Port Perry drew more than 2,500 people from around the world. Auctioneer Bob Shore opened the bidding on Brookview Tony Charity at $50,000. When it stopped, a half-interest in that one cow had brought $1,450,000 — a Canadian record, Stephen B. Roman bidding for Romandale Farms.

That’s what a dispersal was: the moment a breeder’s life’s work got appraised in public, with buyers from three continents leaning on the rail.

Dawdy had bred the sire of Round Oak Rag Apple Elevation. He’d bred Ormsby Burke Pontiac Mose, paternal grandsire of Paclamar Astronaut. His last herd sire was Tidy Burke Cornerstone, a Mose son born in 1957. Any auctioneer in America would have fought for that sale.

He farmed his cows out to a couple of young Kansas breeders he had confidence in, and walked off.

No trumpets. Twenty-four cows quietly handed to two young men he trusted.

Doug Dunton’s ending was quieter still, and to a breeder’s eye it’s the sadder of the two.

He’d built the finest brood cow herd in Canada — and one by one the bulls carrying his blood left. Voyageur. Midnight Joe. Prefect. Radar, Sky Chief, Crown Royal. They finished their careers standing in A.I. studs, working for other people’s catalogs and other people’s prefixes, the same way A.B.C. Inka May’s son had.

Picture Glenvue after they’d gone. The tie-stall rows still full of those matched, deliberate, warm-bred cows. The row of Midnight Joe daughters aging in place. The best young bulls gone to studs whose names would go on the semen straws while the Glenvue prefix quietly stopped appearing at the top of pedigrees.

His best customers — Dawdy’s too — were farmer-breeders who lived no more than twenty miles from the farmgate. That was the distribution network. Neighbors, driving over to look at a young bull.

Twenty miles. For genetics that now circle the planet.

Where they live today — and this part isn’t sentiment; it’s arithmetic

If you breed Holsteins, both of these men are standing in your barn tonight. Not metaphorically.

A Y-chromosome study published in the Journal of Dairy Science traced the paternal pedigrees of 62,897 North American Holstein bulls born between 1950 and 2013 and found the male line had collapsed to essentially two branches.

Among bulls born in the 2010s: Chief’s line at 48.78%. Elevation’s at 51.06%. A third lineage, Penstate Ivanhoe Star, hung on at 0.16% — a whole paternal family down to a rounding error.

Split almost exactly down the middle. Dunton’s bull and Dawdy’s bull, dividing a breed between them.

The Bullvine’s own analysis puts today’s figure at 99.84% of active A.I. sires tracing to those two. Penn State’s Dr. Chad Dechow doesn’t soften it: “When people say 99 percent of Holstein sires in the U.S. today can be traced to only two bulls, both born in the 1960s, it’s absolutely true.”

Chief’s chromosomes account for roughly 14 percent of the entire U.S. Holstein genome. He left about 16,000 daughters, 500,000 granddaughters, and more than two million great-granddaughters.

Elevation sired over 10,000 A.I. sons across 45 countries, with an estimated nine million descendants — and by The Bullvine’s analysis, his DNA still makes up something close to a tenth of the CDCB genomic reference population, the dataset your genomic predictions are trained on.

The pathways run where you’d expect. Chief’s best sons were S-W-D Valiant (June 28, 1973) and Walkway Chief Mark (June 13, 1978, out of Walkway Matt Mamie, a No-Na-Me Fond Matt daughter).

On Elevation’s side: Sweet-Haven Tradition (August 18, 1974) and Hanoverhill Starbuck — born April 26, 1979, out of Anacres Astronaut Ivanhoe, an Astronaut daughter. Which means Dawdy’s Mose line shows up on both sides of Starbuck’s pedigree.

Hanoverhill Starbuck, b. April 26, 1979 — the first bull in this story anybody reading it might have actually used. 200,000-plus daughters across 45 countries. 209 proven sons, 406 proven grandsons. In Canada alone, 36,976 of his daughters got classified, and 70% of them graded Good Plus or better — Doug Dunton’s exact cull rule, cleared at industrial scale by Elmer Dawdy’s bloodline. Read more: Hanoverhill Starbuck’s DNA Dynasty: The Holstein Legend Bridging 20th-Century Breeding to Genomic Futures)

And Starbuck is where you can watch the whole thing come full circle.

He left more than 200,000 daughters across 45 countries and 209 proven sons, which led to 406 proven grandsons. In Canada alone, 36,976 of his daughters were classified — and 70% of them scored Good Plus or better.

Read that number again, because it’s the quiet miracle in this entire story.

Good Plus or better was Doug Dunton’s cull rule. The line he drew in a Brampton tie-stall barn, the standard that decided which cows stayed and which ones went down the road. Seventy percent of thirty-seven thousand daughters of a bull descended from Elmer Dawdy’s overlooked, bought-for-a-song foundation cows would have cleared Doug Dunton’s bar.

The two men never agreed on a bloodline in their lives. Their genetics still met Dunton’s standard anyway.

Ninety-five percent.

One province, one bull, one bloodline.

And in Quebec, 95% of Holsteins carried Starbuck’s genetics. Ninety-five percent — traceable straight back to a five-week-old calf that left Green Bay on a truck in 1947.

Dunton’s Glenvue Noelle Inka threw one more long shadow: her grandson Spring Farm Fond Hope was grandsire of No-Na-Me Fond Matt, and paternal great-grandsire of Hanover-Hill Triple Threat, whose daughter Tora Triple Threat Lulu produced Hanover-Hill Inspiration.

The bill that came due

If this were only a monument, I’d stop there. But these were farmer-breeders, and farmers get the whole ledger.

“You can never get too much of a good thing,” Dawdy said.

He was wrong, and the breed has been paying that invoice ever since.

Chief carried a lethal recessive nobody knew existed — a mutation in the APAF1 gene, now called HH1. One copy makes a carrier. Two copies and there is no calf at all, just a cow that comes back around.

Before it was identified in 2016, that single inherited flaw is estimated to have caused roughly 500,000 spontaneous abortions worldwide and about $420 million in losses across 35 years. Once geneticists found it, carrier frequency fell from 8% to 2% within a few years.

And the concentration those two bulls created is still tightening. Lactanet’s August 2025 figures put average pedigree inbreeding for Canadian Holstein heifers born in 2024 at 9.99%.

Here’s the practical piece, and I think both men would want you to hear it.

That “outcross” bull on your mating list probably isn’t one. When 99.84% of active A.I. sires descend from the same two bulls, a clean-looking four-generation pedigree tells you almost nothing — two animals can show no shared ancestors in three or four generations and still carry a substantial genomic relationship, because the common ancestry sits further back in Chief and Elevation, in both pedigrees, many times over.

That’s what pedigree-based screening can’t see and genomic evaluation can. Run your matings through the CDCB or Lactanet genomic tools rather than reading a paper pedigree and calling it an outcross. The number you want is the genomic inbreeding of the resulting calf, not the count of repeated names on a page.

That’s not a knock on either man. Neither had a sire summary, a daughter proof, or a haplotype screen. They were breeding by eye and by memory. They bred so well, and so consistently, that A.I. took their two best products and made them universal before anybody understood what universal would cost.

The last word

They never met. They’d have argued if they had.

One was a pleasant Ontario dairyman with no degree, two sons who stayed, and a stubborn faith in one bloodline he refused to abandon even once in forty years.

The other was a blunt, degree-holding Kansan who bought crooked cows out of orphanage herds, milked twenty-four of them, traded a fortnight houseguest’s son for a five-week-old calf that became the sire of the century, and had two boys who left and didn’t look back.

Doug Dunton bred the cows. Elmer Dawdy bred the bulls.

E.Y. Morwick spent six pages trying to settle which of them mattered more. He laid out his propositions, ran the pedigrees both directions, weighed Elevation against Chief and Astronaut against Fond Matt, and reached the only honest conclusion available to him: “So far as I can see, they come out pretty well equal.”

Sixty years of genetics have proved him right to two decimal places. 48.78 and 51.06.

And that’s the thing worth carrying out of this story. Some legacies are too large for one man, so history hands them to two — one chasing type when nobody could measure it, one chasing intensity when everybody told him to stop — and lets them build the same thing from opposite ends, in different countries, in barns eleven hundred miles apart, neither one ever knowing the other man was working.

Walk your barn tonight. Look at the udders. Watch how they milk out. Notice which old cow you keep deciding to keep, one more lactation, past what the spreadsheet advises.

Doug Dunton’s three unmeasurable traits are still the ones you’re chasing. Elmer Dawdy’s twenty-four crooked cows are standing in every animal you own.

Two barns. Eleven hundred miles. One breed.

Key Takeaways

  • Neither man had a sire summary, a daughter proof, or a bank account worth mentioning. Dunton had one bloodline he refused to leave; Dawdy had 24 cows nobody wanted and a trade nobody else would make. Scale was never the moat.
  • The cow that founded half the breed’s sire line was classified Good and never raised off it. Before you cull a heifer on her card alone, look at her udder, her mother, and what she throws.
  • Dunton selected for temperament, milk-out, and longevity when none of the three appeared on a pedigree. Seventy years later, that’s the functional-traits conversation the industry finally caught up to.
  • At 99.84% of A.I. sires tracing to Chief and Elevation, a clean four-generation pedigree isn’t proof of an outcross. Run the mating through CDCB or Lactanet and read the calf’s genomic inbreeding instead.

Continue the Story

  • Round Oak Rag Apple Elevation: The Bull of the Century – Step inside the world where Elmer Dawdy’s prison-calf gamble took physical form, tracing how Virginia farmer Ronald Hope guided Elevation from an unheralded young sire into the monumental sire line that conquered modern A.I. studs.
  • Hanoverhill Starbuck – A Legendary Sire – Walk the show barns and test herds where Doug Dunton’s strict type gospel and Dawdy’s linebred genetics finally merged, revealing how one black bull redefined Canadian breeding and proved that the foundation held across generations.
  • Inbreeding in the Dairy Cattle Industry: Is It Time to Panic? – Confront the modern ledger of Chief and Elevation’s absolute reign, unpacking the mounting genetic bottleneck, lethal recessives, and why today’s commercial breeders are navigating the costly consequences of two men who bred too well.

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99.84% of Holstein AI Bulls Trace to Just Two Fathers

Two bulls born in the 1960s—Chief and Elevation—sit behind 99.84% of today’s AI sires. The gift: more milk, better udders. The bill: a 9.99% inbreeding tab now in your heifer pen.

Picture every Holstein in North America walking into one barn for Father’s Day dinner.

Millions of black-and-white cows, shoulder to shoulder, in a building the size of a county. They’ve come to toast their fathers, the way families do this time of year. And here’s the part that ought to stop you cold while you’re scraping the parlor this Sunday: almost every animal in that impossible room would be raising a glass to the same two dads.

Not two dozen. Not two hundred. Two.

Their names were Pawnee Farm Arlinda Chief and Round Oak Rag Apple Elevation. One arrived on an Indiana spring morning in 1962. The other showed up in 1965, on a modest Virginia farm nobody had heard of. Neither ever knew the other. And yet a Y-chromosome study that combed through 62,897 bulls born between 1950 and 2013 found that virtually every active North American Holstein AI bull traces its paternal line back to just these two grandfathers. The Bullvine’s own analysis of that work puts the figure at 99.84% of active AI bulls — split almost eerily down the middle, roughly half Chief and half Elevation. 

Read that number again. Ninety-nine point eight four percent. It’s as if the entire breed flipped a coin sixty years ago and has been living with the result ever since.

So, before you pour your coffee and head out to check the fresh pen, let me properly introduce you to the two dads at the head of your herd’s table. Once you know their story, you’ll never look at your milking string the same way again.

The $4,300 gamble that started a dynasty

Pawnee Farm Arlinda Chief (1962–1982). The bull behind half the breed. From a dam who sold for $4,300, Chief sired 16,000 daughters and more than two million great-granddaughters—and carried a hidden HH1 recessive that the breed wouldn’t decode for fifty years. Read more: The $4,300 Gamble That Reshaped Global Dairy Industry: The Pawnee Farm Arlinda Chief Story

Start with the elder. In a family reunion, you always start with the elder.

Here’s the thing about Chief, though — the gamble that made him happened before he ever drew breath. At the Pawnee Farm dispersal, his dam, Pawnee Farm Glenvue Beauty (EX-90), crossed the auction block and sold for $4,300. Now, picture what that meant in 1962. You could buy a new car twice over. You could put serious money down on land. Somebody stood at that ring, looked at a cow, and decided she was worth more than a house lot. 

They turned out to be right in a way nobody could have predicted.

Because Beauty’s son became a kind of one-animal continent. By the time the dust settled, Chief had produced 16,000 daughters, 500,000 granddaughters, and more than two million great-granddaughters. Stack that against the cow your grandfather was proud to own, and you start to feel the gravity of the thing. This wasn’t a good bull. This was a whole population’s worth of fatherhood compressed into one animal. 

And his daughters could milk. The proof has a name — Beecher Arlinda Ellen. In 1975, on Harold Beecher’s farm near Rochester, Indiana, Ellen completed a lactation of 55,661 pounds, the first cow in the entire Holstein breed to crack 55,000 in a single year — a world record that would stand for nearly two decades.

Here’s the part worth sitting with. Reporters came calling, the way they do when a farm makes history, and asked what magic ration he’d been feeding her. By Harold Beecher’s own account, he hadn’t done anything special at all. Think about that for a second. A humble Indiana dairyman, a world record standing in his tie-stall, and his honest answer was a shrug. He knew what every good cowman knows — you don’t feed your way to a number like that. You breed your way there. Ellen wasn’t a fluke. She was Chief’s signature, written in the milk tank.

And here’s what made Chief’s story the hard one. This was the era before genomics — no DNA test to whisper which young bull was worth sampling. You bred him, you waited, and you milked his daughters for years before the herd finally told you whether you were holding a fortune or a flop. Chief’s people waited. And the daughters kept coming back with the same verdict, herd after herd, in barns that had never heard of Pawnee Farm: more milk, again, and again. A father proves himself slowly. Chief proved himself the only way the times allowed — and the breed was never the same after the proof came in.

When sons become legends in their own right

Great fathers don’t just have great children. They have children who become great fathers themselves — and that’s where Chief’s story gets bigger than one bull.

S-W-D Valiant (EX-95 GM). One of Chief’s great sons. Born June 28, 1973, out of a VG-85 Admiral dam, Valiant took his father’s milk and added show-ring type—the kind of son who becomes a legend in his own right and keeps the family table growing. Read more: The S-W-D Valiant Story: How Genetics Promised Everything and Changed How We Think About Breeding

His most influential sons read like a roll call: Walkway Chief Mark, S-W-D Valiant, Glendell Arlinda Chief, and Milu Betty Ivanhoe Chief. Take Walkway Chief Mark. He was only ever sampled because his full brother died, and somebody needed a backup. The spare. That backup bull accounted for roughly 7% of every Holstein genome on this continent. (The Bullvine has told that whole strange, wonderful story in full in Walkway Chief Mark’s profile — it’s one of the great accidents in breeding history.) 

Seven percent. From the understudy.

Walkway Chief Mark (VG-87 GM). The spare that ran the breed. Only sampled because his full brother died and Foster Walk’s Illinois herd needed a backup, Mark went on to account for roughly 7% of every Holstein genome in North America. Select Sires later named him an Impact Sire of the Breed. The understudy nobody saw coming. Photo: Remsberg. Read more: Walkway Chief Mark: The Backup Bull Behind Seven Percent of Every Holstein Cow

A powerful father’s influence doesn’t stop with his own kids. It compounds. It ripples down through sons, and their sons, until you can’t open a modern catalog without bumping into the old man’s name a dozen times over. Chief didn’t just have a big family. He had a big family that kept having big families, branch after branch — one line eventually threading down to To-Mar Blackstar, himself one of the most heavily used bulls in breed history. Generation after generation, the table just kept getting longer, and the gambler who paid $4,300 for a cow back in 1962 kept looking smarter. 

To-Mar Blackstar. The branch that kept growing. Down one of Chief’s many lines, Blackstar became one of the most heavily used bulls in breed history—proof of how a great father’s influence doesn’t stop with his sons, but compounds, generation after generation, until you can’t open a catalog without bumping into the old man’s name. Photo: Remsberg. Read more: To-Mar Blackstar: The One-Embryo Holstein Sire Behind 15.8% of Today’s DNA – and the Genetic Debt in Your Herd

The B-team mating that produced the Bull of the Century

Round Oak Rag Apple Elevation (1965–1979). The Bull of the Century. He came from a fertility-troubled sire and a “B-team” dam nobody expected anything from—then sired over 10,000 AI sons across 45 countries and an estimated nine million descendants worldwide. The cousin’s hunch that built the barns at Select Sires. Photo: Remsberg. Read more: Round Oak Rag Apple Elevation: The Bull That Changed Everything

Now, meet the other grandfather. And get ready to be surprised, because Elevation’s beginning was the opposite of a sure thing.

Down on Round Oak Farm in the Virginia piedmont, Ronald A. Hope and his family were running a working dairy, not a genetics empire — the kind of modest operation you’d have driven past a hundred times without a second look. The mating that produced Elevation wasn’t some master plan off a proof sheet. It came from Ron Hope’s cousin, George Miller, who suggested the cross. Just a hunch, passed down within the family. Try this one. 

And honestly, on paper, you’d have shrugged and moved on. The sire, Tidy Burke Elevation, had fertility trouble. The dam, Round Oak Ivanhoe Eve, had been shuffled onto the farm’s B-team because she matured too slowly. A questionable father. An overlooked mother. A cousin’s offhand suggestion.

What walked out of that barn in 1965 changed the world. 

I don’t say that lightly. He would later be named the “Bull of the Century.” But forget the title for a second and walk into a parlor full of his daughters instead. Look up. There it is — the udder. High, wide, held tight to the body, still bolted on the way you’d want it two and three lactations after the cows around it had broken down and shipped. Watch one of those daughters walk: sound on her feet into her sixth lactation, settling back in calf as if it were nothing, walking up to milk at an age when her contemporaries were long gone. That’s what Elevation transmitted — and the remarkable thing is he transmitted it all at once: production, udder quality, mobility, fertility, and longevity, in one package, when breeders had spent generations trading one good trait away to get another.

Put numbers on it, and your cup goes down on the table. His daughters averaged about 29,500 pounds in their first lactation — roughly 15% above their contemporaries in the 1970s. And while the industry average was near 2.8 lactations per cow, Elevation’s daughters averaged 4.2. 

Do the barn math on that. Your average cow leaves after 2.8 lactations. Your neighbor’s Elevation daughters are still walking into the parlor at 4.2. Same feed bill to raise the heifer, same calving, and he’s getting roughly half again the productive life out of every replacement. That’s not a show-ring statistic. That’s a mortgage payment. For the farmer living it, the whole thing came down to a simple difference: a cow you fought all year, versus one you forgot to worry about.

A father whose children fill 45 countries

If Chief built his dynasty through a few towering sons, Elevation built his through sheer abundance.

Over 10,000 of his sons became registered AI sires. His semen was shipped to 45 countries. And his descendants — brace yourself — run an estimated 8.8 to 9 million worldwide. There are whole nations with fewer people than this one bull has grandchildren. 

Hanoverhill Starbuck (1979–1998). Elevation’s most famous son. A $2,500 calf whose semen would eventually sell for roughly $25 million, Starbuck sired over 200,000 daughters across 45 countries—and by the early 2000s, some 93% of Canadian Holsteins traced back to him. The Canadian Holstein Association called him, simply, “the Best.” Shown here at five. Photo: Jim Rose. Read more: Hanoverhill Starbuck’s DNA Dynasty: The Holstein Legend Bridging 20th-Century Breeding to Genomic Futures

The most famous of those children crossed the border into Canada and became a legend in his own right: Hanoverhill Starbuck, a $2,500 calf whose semen eventually sold for roughly $25 million. (Starbuck’s story deserves its own evening — The Bullvine has told it in full.) Through Starbuck and ten thousand other sons, Elevation became the patriarch at the head of dinner tables from Wisconsin to the Netherlands to Japan. 

Johanna Rag Apple Pabst, Grand Champion, mid-1920s. Where the family tree begins. The “Rag Apple” buried in Chief’s name and the bloodline behind Elevation’s dam both run back to this one Wisconsin bull—undefeated in 1924 and the foundation ancestor whose name still rides in pedigrees a century on. Walk far enough up the tree, and both grandfathers shake hands here. Read more: The Bull Who Changed Everything: The Johanna Rag Apple Pabst Story

And here’s a detail that ties the whole tree together. Eve — the overlooked B-team mother nobody expected anything from — traced back twenty times to a foundation cow named Johanna Rag Apple Pabst. The “Rag Apple” buried in Chief’s name comes from the same deep well. These two grandfathers, born to different farms in different decades, weren’t strangers at all. Walk far enough up the family tree, and they shake hands. The reunion was always a family affair. 

Northcroft Ella Elevation (EX-97 4E GMD DOM). Both grandfathers in one cow. Born February 26, 1974, Ella carried Elevation on top and an EX-91 Chief daughter underneath—the two bloodlines that fathered half the breed, shaking hands in a single pedigree. The reunion, made flesh. Photo: Remsberg.

Two fathers, two temperaments

Set the two old bulls down at the same table, and you’d have spotted the difference fast. They were nothing alike.

Chief was the quiet workhorse — a production sire whose genius announced itself in the milk tank, lactation after lactation, value measured in pounds and years rather than ribbons. Elevation was the showman with substance, one of the first proven bulls of the modern era who could put a daughter in the ring and fill the bulk tank. One made cows that paid. The other made cows that paid and turned heads on the colored shavings.

Elevation did something else, too — he changed the very machinery that moves genetics around the world. His semen, by one account, helped finance Select Sires and solidify it as a cooperative during its fragile early years. As his own breeder’s cousin, George Miller, put it: “It’s been said that Elevation built the barns at Sire Power and Select Sires.” 

And his fingerprints are still all over the modern toolbox. Here’s the mind-bender: by The Bullvine’s analysis, Elevation’s DNA makes up about 8.3% of the CDCB’s genomic reference population — the very dataset that modern genomic predictions are trained on. Think about that the next time a young genomic bull’s numbers flash up on your screen. The math ranking him was partly based on his own great-great-grandfather. 

The roots run deeper than you think

Speaking of walking up the tree, the story doesn’t actually start with these two.

Dr. Chad Dechow’s work shows that all the great 1960s pillars of the breed trace their male lines back to just two bulls born in the early 1880s: one called Neptune H, born in 1880, and one named Hulleman, born in 1881.

Sit with that. The Father’s Day table you’ve been picturing doesn’t have two chairs at the head — it has two chairs in this generation. Keep walking back, and the whole enormous family narrows again and again until, in the 1880s, it comes down to a pair of bulls who lived before the automobile, before the milking machine, before electricity reached most farms.

We like to think we’re steering. Our index, our matings, our careful selection — surely that puts us in the driver’s seat. And it does, a little. But we’re steering a river that’s been running in the same channel for nearly 140 years. Someday, a breeder none of us will ever meet will trace a herd back to a bull you used this week, and they’ll feel exactly the way you feel as you read these names right now. That’s the strange gift of a breed this old. You’re never just raising cattle. You’re handing something down.

The morning the numbers didn’t add up

Now comes the hard part of every honest Father’s Day — the part where you love somebody and still have to tell the truth about them.

It started, in a way, with researchers staring at a spreadsheet that made no sense.

In 2011, USDA scientists were studying haplotypes — long stretches of chromosome inherited as a single block — when they noticed something wrong on chromosome 5. A particular haplotype was common across the breed. Carriers were everywhere. By the plain arithmetic of inheritance, there should have been thousands of living animals carrying two copies of it. They went looking for those animals. There were none. Not a single one. The double-carriers weren’t dying young or growing up sickly — they were never being born at all. 

Five years later, a team led by Heather Adams with USDA’s Paul VanRaden ran the cause to ground: a single “nonsense” mutation in a gene called APAF1, a typo that truncates more than half the protein it’s supposed to build. 

One copy, and a calf is just a carrier — perfectly healthy. But breed a carrier to a carrier, which is heartbreakingly easy when half the breed descends from the same grandfather, and two copies quietly kill the embryo before it’s ever born.

They traced the haplotype straight back to Chief. And before anyone knew it was there, that single inherited flaw is estimated to have caused roughly half a million spontaneous abortions worldwide — and about $420 million in losses over 35 years. The flip side runs staggeringly in the other direction: the same researchers estimate that Chief’s beneficial genetics added about $30 billion in increased milk production. The gift and the bill, written into the same animal. 

Half a million calves conceived and quietly lost. Half a million heat checks that came up empty — a farmer standing in the barn at dusk, wondering what went wrong, never knowing the answer had been written into the breed’s most celebrated father sixty years before he was born.

That’s no reason to resent Chief. A father doesn’t choose the genes he carries. But it’s the unavoidable math of a narrow family tree: when everyone shares the same grandfather, his hidden flaws stop being rare. The good news — once the mutation had a name, breeders could test for it and breed around it, and U.S. carrier frequency fell from roughly 8% to about 2% within a few years. The defect didn’t end Chief’s legacy. It just made us smarter about how we carry it forward. 

The number landing in your heifer pen right now

Here’s where the history stops being history.

According to Lactanet’s August 2025 update, the average pedigree-based inbreeding of Canadian Holstein heifers born in 2024 hit 9.99%. Nearly ten percent. A generation ago, that figure would have set off alarms. Today it’s just Tuesday. 

That’s what two grandfathers at the head of the table eventually costs a family. Every percentage point of inbreeding chips away at fertility, at calf vigor, at the very longevity that made Elevation famous in the first place. The traits these great fathers gave us are exactly the ones a too-narrow pedigree slowly takes back. By the USDA’s measure, Dr. Dechow puts both Chief and Elevation at a genetic relationship of about 14% to the modern Holstein cow. Two bulls, wearing different hides, make up a huge chunk of your herd. (The Bullvine has run the dollars-and-cents of where this is heading in its breakdown of Holstein’s inbreeding bill.) 

Maxima de Bois Seigneur. Sixty years later, still in the room. A daughter of Stantons Chief—and a direct descendant of Pawnee Farm Arlinda Chief—Maxima stands in a Belgian farmyard as living proof that the old grandfather never left the table. Every time you see a modern cow like her, you’re looking at his influence. Photo: Guillaume Moy. Read more: From Laurie Sheik to Robotic Milking: Bois Seigneur Holstein’s Journey of Innovation

What this means for your operation

Here’s the good news in all of this: knowing the family history is exactly what lets you manage it. So you’ve met the grandfathers — what do you actually do with this on Monday morning? A few concrete things.

Run your matings through a genomic inbreeding tool, not just a pedigree check. With 99.84% of AI sires tracing to two bulls, pedigree alone hides how related your “outcross” really is. The genomic future inbreeding value tells the truth.

Check carrier status for HH1 (APAF1) before you breed a deep-Chief cow. Most catalogs list it. Avoiding carrier-to-carrier matings is the cheapest insurance you’ll ever buy against an empty calving pen.

Put a hard ceiling on expected progeny inbreeding. Many breeders aim to keep a mating under roughly 6–7%. With the Canadian average heifer already at 9.99%, every mating you pull below that line is a small win for the next generation.

Actively hunt the rare outcross lines. They exist. They’re harder to find, and they’re worth the search — the breed’s long-term fertility depends on the breeders who refuse to let the family tree narrow any further.

None of this is a knock on Chief or Elevation. You’d have made the same call any of those old breeders made — the production was real, the longevity was real, the money was real. This is simply the next chapter of stewardship: honoring what the grandfathers built while quietly widening the table for everyone who comes after.

The reunion, and what we owe the dads at the table

Come back to that impossible barn one last time.

The millions of cows. The two chairs at the head. The two old bulls who never met and yet fathered nearly all of it — one a $4,300 gamble out of Pawnee Farm, one a cousin’s hunch off a modest Virginia hillside that had no business working and changed everything anyway. Between them, they handed the dairy world more milk, better udders, longer-lasting cows, and a uniformity that built the modern industry. They also handed down a narrower gene pool and a few hidden flaws their children are still reckoning with. Both things are true. That’s what it means to inherit from a great father — the gifts and the burdens come in the same package, and the work of a lifetime is sorting out what to do with each.

So this Sunday, when somebody asks what you do for a living, tell them the truth. You’re raising the great-great-grandchildren of two bulls born in the 1960s — who themselves came down from a pair born in the 1880s — in a family reunion that has never once adjourned, and never will.

Pour a little extra in the cup. The grandfathers earned it.

Key Takeaways

  • That “outcross” bull on your mating list probably isn’t one — 99.84% of active AI sires trace to Chief or Elevation, so run matings through a genomic inbreeding tool, not just the pedigree.
  • Before you breed a deep-Chief cow, check HH1 (APAF1) carrier status on both sides; a carrier-to-carrier mating is the cheapest way to end up with an empty calving pen.
  • The traits these two gave us — milk, udders, longevity — are the same ones a narrow pedigree quietly takes back, so aim to keep expected progeny inbreeding under roughly 6–7%, against a breed-average heifer already at 9.99%. 
  • The breed’s long-term fertility depends on the breeders who hunt and use the rare outcross lines — they’re harder to find, and they’re worth the search. 

Methodology Note

This article uses several distinct measures of genetic influence that should not be conflated. The 99.84% figure is a paternal Y-chromosome lineage measure derived from Yue et al. (2015, Journal of Dairy Science 98(4):2738–2745, examining 62,897 bulls) — it describes male-line descent, not total genome share; the 99.84% / roughly-half-each breakdown is The Bullvine’s analysis of that dataset. The genetic relationship to the modern Holstein cow (~14% for both bulls) comes from Dr. Chad Dechow’s USDA-affiliated analysis, as reported in Hoard’s Dairyman. The Bullvine reports that Elevation accounts for approximately 8.3% of the CDCB genomic reference population. The HH1/APAF1 facts come from the 2011 USDA haplotype discovery (VanRaden et al., J. Dairy Sci. 94:6153–6161) and Adams et al. (2016, J. Dairy Sci. 99(8):6693–6701), which identified the causative APAF1 nonsense mutation. The estimates of roughly half a million abortions, about $420 million in losses over 35 years, and about $30 billion in beneficial milk production are reported by UC Davis (2016). The 9.99% inbreeding figure is a pedigree-based coefficient for Canadian Holstein heifers born in 2024 (Lactanet, August 2025) and may differ from U.S. CDCB genomic measures. National figures may not reflect your region or herd; verify carrier status and inbreeding values against current CDCB/Lactanet data for your own matings.

Questions, corrections, or a number you’d like us to double-check? Reach out to editor@thebullvine.com

Continue the Story

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Round Oak Rag Apple Elevation: The Bull That Changed Everything

Born from an unlikely mating, one bull revolutionized global dairy breeding with genetics so powerful they still dominate herds 60 years later.

Round Oak Rag Apple Elevation (1965-1979), the legendary Holstein sire dubbed “Bull of the Century,” photographed in his prime at Select Sires. This unassuming black and white bull from Virginia transformed global dairy genetics with his exceptional ability to transmit production, conformation, and longevity traits simultaneously. Note his balanced frame, strong topline, and characteristic Elevation profile—physical traits that would be passed to over 8.8 million descendants worldwide. While unremarkable by today’s extreme standards, this bull’s genetic blueprint revolutionized Holstein breeding and continues to influence elite dairy cattle six decades later. His balanced genetics remain the gold standard for functional type: not too tall, not too extreme, but built to last. Photo: Remsberg.
Round Oak Rag Apple Elevation (1965-1979), the legendary Holstein sire dubbed “Bull of the Century,” photographed in his prime at Select Sires. This unassuming black and white bull from Virginia transformed global dairy genetics with his exceptional ability to transmit production, conformation, and longevity traits simultaneously. Note his balanced frame, strong topline, and characteristic Elevation profile—physical traits that would be passed to over 8.8 million descendants worldwide. While unremarkable by today’s extreme standards, this bull’s genetic blueprint revolutionized Holstein breeding and continues to influence elite dairy cattle six decades later. His balanced genetics remain the gold standard for functional type: not too tall, not too extreme, but built to last. Photo: Remsberg.

Do you know how some legends never fade? Well, in the dairy world, there’s one name that still makes breeders sit up straighter when mentioned – Round Oak Rag Apple Elevation. I can’t tell you how often I’ve heard old-timers at cattle shows talk about this bull with a reverence usually reserved for religious figures. And honestly, they’re not wrong.

Born in 1965 on a modest Virginia farm, this unassuming black-and-white calf would become the most significant genetic influencer Holstein breeding has ever seen. Can you believe his bloodline now runs through nearly 9 million descendants? That’s right – almost every glass of milk you’ve ever enjoyed likely came from a cow with some connection to this legendary sire.

What made Elevation so special? He delivered both, unlike most bulls that give you either production OR pretty cows. His daughters pumped an incredible 29,500 pounds of milk during their first lactations – beating their peers by 15%! – while sporting those picture-perfect udders that look like they were crafted by a sculptor with an obsession for symmetry. You’ve gotta appreciate a bull that refuses to compromise.

I’ve always found it fascinating that his story began with what you might call a questionable mating. His sire had fertility issues, and his dam was considered too slow-maturing for the fast-paced dairy world. This pairing might never have happened in today’s era of genomic testing and algorithm-driven breeding programs. Kinda makes you wonder what other genetic gold mines we’re potentially missing by being too reliant on numbers.

The Unlikely Star: How Elevation Came to Be

George Miller, a pivotal figure in Holstein breeding history and cousin to Ronald Hope Sr., photographed during his tenure as marketing manager at Select Sires. Growing up on his uncle’s Round Oak Farm in Virginia, Miller was instrumental in planning the legendary mating that produced Round Oak Rag Apple Elevation. After earning degrees in dairy science from Virginia Polytechnic Institute, Miller’s career spanned from managing Virginia Artificial Breeders Association to joining Select Sires in 1973, where former general manager Dick Chichester praised him as “honest” and committed to “doing things right.” Miller’s passionate advocacy for balanced genetics and his foresight regarding ELEVATION’s potential helped transform dairy breeding practices worldwide. Following his passing in February 2021 at age 94, Select Sires established the George Miller Memorial Scholarship Fund to honor his legacy. Photo courtesy of Select Sires Archives.
George Miller, a pivotal figure in Holstein breeding history and cousin to Ronald Hope Sr., photographed during his tenure as marketing manager at Select Sires. Growing up on his uncle’s Round Oak Farm in Virginia, Miller was instrumental in planning the legendary mating that produced Round Oak Rag Apple Elevation. After earning degrees in dairy science from Virginia Polytechnic Institute, Miller’s career spanned from managing Virginia Artificial Breeders Association to joining Select Sires in 1973, where former general manager Dick Chichester praised him as “honest” and committed to “doing things right.” Miller’s passionate advocacy for balanced genetics and his foresight regarding ELEVATION’s potential helped transform dairy breeding practices worldwide. Following his passing in February 2021 at age 94, Select Sires established the George Miller Memorial Scholarship Fund to honor his legacy. Photo courtesy of Select Sires Archives.

So here’s the backstory that sounds more like a feel-good movie than real life. Two cousins, Ronald Hope Sr. and George Miller, had spent a quarter-century meticulously layering Burke and Ivanhoe bloodlines into their herd at Round Oak Farm. Talk about playing the long game! These guys weren’t chasing quick wins but building something meant to last.

In 1965, they made a decision that probably raised some eyebrows. They bred Tidy Burke Elevation (a bull with questionable fertility) to Round Oak Ivanhoe Eve (a cow relegated to the farm‘s “B-team” because she matured too slowly). It wasn’t exactly a match made in bovine heaven, at least on paper.

But man, did that gamble pay off! This unlikely coupling created a genetic alchemy that would transform dairy farming forever. Elevation inherited the milk-producing magic from his sire’s Burke lineage while getting structural soundness and mammary excellence from his dam’s Ivanhoe connections. The result? A genetic unicorn whose DNA contained a rare chromosome 6 haplotype that optimized fat-to-protein ratios – something we didn’t even have the technology to identify until decades later!

You might be surprised to learn that Elevation wasn’t an instant sensation. His progeny consistently came in second place at early shows – never first. Judges didn’t know what to make of his balanced but unspectacular offspring. They weren’t the tallest, broadest, or most extreme in any category. They were just… good at everything. Talk about being ahead of your time!

Ironically and fitting, this “runner-up” status would eventually prove prophetic. While more specialized, flashier bloodlines came and went, Elevation’s descendants ultimately dominated milking parlors and show rings worldwide. Their versatility outlasted everything else.

Five Qualities That Made Elevation a Legend

If you’ve spent time around dairy farms, you know that most bulls have one standout trait – maybe great production or fancy udders. But Elevation? This guy was the complete package. He redefined Holstein’s breeding through five key characteristics that created what I like to call the “genetic royal flush.”

First up: production. His daughters weren’t just good milkers – they were milk-making machines. Averaging 29,500 pounds in their first lactations during the 1970s put them 15% ahead of their contemporaries. And unlike bulls that give you quantity at the expense of quality, Elevation’s daughters maintained excellent butterfat and protein percentages. You couldn’t ask for more!

Then there were those udders – my goodness, those udders! Charlie Will, who knew these cows better than most, described them as having “high and wide rear udders with exceptional shape and symmetry.” This wasn’t just pretty – it was functional. These udders stayed attached 2-3 lactations longer than average, translating to an extra $1,200 profit per cow back in the 70s. Not too shabby!

Mobility might not be the sexiest trait to discuss over coffee, but it’s a game-changer on the farm. Elevation’s girls showed up with “straight legs, healthy hocks, and strong loins” – dull on paper, maybe, but pure gold in practice. These cows stayed sound even on concrete floors (the bane of many dairy cows’ existence), allowing them to keep producing at high levels for 5-7 lactations when most cows were burning out after 3.

Fertility might be the most underrated of Elevation’s gifts. His daughters rebred 14 days faster than their herd mates – two weeks might not sound like much, but multiply that across thousands of cows and millions of lactations, and you’re talking serious money. Plus, this reproductive efficiency helped his genetics spread like wildfire.

Finally, there’s longevity – the crown jewel. While the industry average was 2.8 lactations per cow in the 1970s, Elevation’s daughters stuck around for 4.2. That’s a 50% increase in productive life! Herds with his bloodlines reported 22% lower replacement costs. For farmers operating on tight margins, this was revolutionary.

What sets Elevation apart wasn’t excelling in any category – it was his “genetic coherence,” the ability to transmit ALL these qualities simultaneously without trade-offs. It’s like getting a sports car with excellent gas mileage or a delicious and healthy dessert. Usually, you don’t get both, but with Elevation, you did!

OLMAR ELEVATION DAZZLING STAR (EX-94 GMD), photographed in her prime, exemplifies the exceptional type and production balance that made Elevation daughters legendary. Her strong, well-attached mammary system and correct dairy structure showcase the genetic superiority that earned her both an Excellent classification and Gold Medal Dam status—hallmark achievements reflecting Elevation’s ability to produce daughters who excelled in both the show ring and milking parlor. Note her combination of dairy strength, angularity, and impressive udder capacity—traits that contributed to extended productive life and the “genetic coherence” discussed in our article. Photo: Pete’s Photo
OLMAR ELEVATION DAZZLING STAR (EX-94 GMD), photographed in her prime, exemplifies the exceptional type and production balance that made Elevation daughters legendary. Her strong, well-attached mammary system and correct dairy structure showcase the genetic superiority that earned her both an Excellent classification and Gold Medal Dam status—hallmark achievements reflecting Elevation’s ability to produce daughters who excelled in both the show ring and milking parlor. Note her combination of dairy strength, angularity, and impressive udder capacity—traits that contributed to extended productive life and the “genetic coherence” discussed in our article. Photo: Pete’s Photo

How One Bull Transformed an Industry

You know what’s crazy? Elevation didn’t just change individual herds—it reshaped entire organizations and industry practices. I’ve talked with folks who worked at Select Sires during that era, and they’ll tell you straight up: “Elevation put Select Sires on the map.”

In the ’60s, Select Sires struggled to establish itself as a newly formed federation of regional breeding organizations. Then this bull came with his perfect combination of production and type, and suddenly, everyone wanted Select Sires’ genetics. The revenue from Elevation semen sales built the company’s infrastructure. George Miller said it best: “It’s been said that Elevation built the barns at Sire Power and Select Sires.”

Think about that impact for a minute. One Bull’s genetics were so sought-after that they funded buildings, grew market share, created brand identity, and helped merge 18 state-level organizations into a cohesive national presence. That’s not just breeding success – that’s business transformation!

His influence spread well beyond American borders, too. Elevation’s semen was shipped to 45 countries, fundamentally reshaping global Holstein breeding. He served as a Holstein ambassador, making friends for American genetics worldwide. In Canada, his impact was especially pronounced through his son Hanoverhill Starbuck, who became the cornerstone of Canadian breeding programs. European dairy industries in France, Italy, and the Netherlands incorporated his bloodlines to improve their national herds. Elevation descendants eventually made up 70% of the Holstein population in some countries, like France!

Developing dairy nations used Elevation genetics to rapidly modernize their herds, while emerging dairy industries in Asia used their bloodlines to establish foundation herds adapted to local conditions. He created a genetic standardization that connected Holstein populations worldwide –bovine globalization, if you will!

A poignant moment in dairy breeding history: Mr. and Mrs. Ronald Hope of Purcelville, Virginia (far left) receive a painted portrait of Round Oak Rag Apple Elevation at his memorial dedication ceremony on August 1, 1979. The group stands behind Elevation’s permanent gravestone at Select Sires headquarters, which marks his life from August 30, 1965, to April 25, 1979. The ceremony honored the bull that transformed the Holstein breed and financially secured Select Sires’ future. Also pictured are Robert Rumler of Holstein Association, Dick Chichester and George Miller of Select Sires—the latter being Hope’s cousin who suggested the legendary mating that produced Elevation. The memorial site, positioned by the reflecting pond at Select Sires’ main entrance, remains a pilgrimage destination for dairy breeding enthusiasts worldwide. Photo: Johnson/Select Sires Archives.
A poignant moment in dairy breeding history: Mr. and Mrs. Ronald Hope of Purcelville, Virginia (far left) receive a painted portrait of Round Oak Rag Apple Elevation at his memorial dedication ceremony on August 1, 1979. The group stands behind Elevation’s permanent gravestone at Select Sires headquarters, which marks his life from August 30, 1965, to April 25, 1979. The ceremony honored the bull that transformed the Holstein breed and financially secured Select Sires’ future. Also pictured are Robert Rumler of Holstein Association, Dick Chichester and George Miller of Select Sires—the latter being Hope’s cousin who suggested the legendary mating that produced Elevation. The memorial site, positioned by the reflecting pond at Select Sires’ main entrance, remains a pilgrimage destination for dairy breeding enthusiasts worldwide. Photo: Johnson/Select Sires Archives.

The Family Tree That Changed Everything

Want to know what cemented Elevation’s legacy? His sons – over 10,000 of them became registered AI sires! That’s an army of genetic influence that’s almost impossible to comprehend. Some of his most influential sons include Sweet-Haven Tradition, Rockalli Son of Bova, Marshfield Elevation Tony, Ocean-View Sexation, and Straight-Pine Elevation Pete. Charlie Will also highlights Mars Tony and Lime Hollow Mars as influential Elevation sons.

Straight-Pine Elevation Pete, one of Round Oak Rag Apple Elevation’s most influential sons, photographed in the early 1970s. Pete carried forward his sire’s exceptional genetic traits for production and conformation, helping to establish the Elevation bloodline throughout North American Holstein herds. Note his balanced frame, strong topline, and dairy character—hallmarks of the structural soundness that made Elevation progeny legendary for their longevity and productive life. Photo credit: Remsberg.
Straight-Pine Elevation Pete, one of Round Oak Rag Apple Elevation’s most influential sons, photographed in the early 1970s. Pete carried forward his sire’s exceptional genetic traits for production and conformation, helping to establish the Elevation bloodline throughout North American Holstein herds. Note his balanced frame, strong topline, and dairy character—hallmarks of the structural soundness that made Elevation progeny legendary for their longevity and productive life. Photo credit: Remsberg.

But if we’re talking about Elevation’s sons, we’ve got to spotlight Hanoverhill Starbucks. If Elevation were the king, Starbucks would have been the crown prince who expanded the dynasty. His impact on global Holstein genetics was profound, especially in Canada. Starbucks sons like Madawaska Aerostar, Besne Buck, Fatal, and Sabbiona Bookie carried Elevation’s genes into another generation with even more significant influence.

Hanoverhill Starbuck, one of the most influential Holstein sires in dairy history, captured here at 5 years old by photographer Jim Rose. Standing an impressive 73½ inches at the shoulder (1.87m) and weighing 2,580 lbs (1,173 kg), Starbuck’s exceptional feet and leg quality—evident in this profile—became his trademark and a key factor in his global genetic impact. Born in 1979 and sired by the legendary Round Oak Rag Apple Elevation, Starbuck went on to father over 200,000 daughters and 209 proven sons across 45 countries, earning the “Premier Sire” title 27 times at major Holstein shows. His balanced frame, perfect leg set, and overall structural correctness revolutionized Holstein breeding, with an estimated 93% of Canadian Holsteins born between 2003-2005 tracing back to this remarkable bull. The Canadian Holstein Association eventually dubbed him “Simply the Best”—a title that begins to explain why his genetics remain influential in dairy herds worldwide nearly three decades after his passing in 1998. Photo: Jim Rose.
Hanoverhill Starbuck, one of the most influential Holstein sires in dairy history, captured here at 5 years old by photographer Jim Rose. Standing an impressive 73½ inches at the shoulder (1.87m) and weighing 2,580 lbs (1,173 kg), Starbuck’s exceptional feet and leg quality—evident in this profile—became his trademark and a key factor in his global genetic impact. Born in 1979 and sired by the legendary Round Oak Rag Apple Elevation, Starbuck went on to father over 200,000 daughters and 209 proven sons across 45 countries, earning the “Premier Sire” title 27 times at major Holstein shows. His balanced frame, perfect leg set, and overall structural correctness revolutionized Holstein breeding, with an estimated 93% of Canadian Holsteins born between 2003-2005 tracing back to this remarkable bull. The Canadian Holstein Association eventually dubbed him “Simply the Best”—a title that begins to explain why his genetics remain influential in dairy herds worldwide nearly three decades after his passing in 1998. Photo: Jim Rose.

The Starbucks line shows the power of Elevation’s genetics—they didn’t dilute over generations; they often amplified! Madawaska Aerostar became one of the first bulls to sell one million doses of frozen semen. His sons in Canada included Maughlin Storm and the Millionaire Sires Startmore Rudolph and Oliveholme Aeroline. Meanwhile, Besne Buck’s son, Jocko Besn, became so influential in France that he sired more than 50% of French Holstein cattle!

Northcroft Ella Elevation (EX-97 GMD DOM), one of Round Oak Rag Apple Elevation’s most celebrated daughters, photographed in her prime. Born February 26, 1974, Ella exemplifies the genetic perfection that made Elevation’s progeny legendary. Her flawless dairy structure, exceptional mammary system, and balanced frame earned her Supreme Champion honors at World Dairy Expo in 1980, where judges declared her “the new ideal Holstein cow.” This breeding masterpiece—out of an EX-91 GMD DOM Pawnee Farm Arlinda Chief dam—later received All-Time All-American Aged Cow recognition in 1984, cementing her place among the greatest show cows in Holstein history. Ella represents the pinnacle of Elevation’s ability to transmit both exceptional type and production traits simultaneously, embodying the “genetic coherence” that made her sire the Bull of the Century. Photo credit: Jack Remsberg.
Northcroft Ella Elevation (EX-97 GMD DOM), one of Round Oak Rag Apple Elevation’s most celebrated daughters, photographed in her prime. Born February 26, 1974, Ella exemplifies the genetic perfection that made Elevation’s progeny legendary. Her flawless dairy structure, exceptional mammary system, and balanced frame earned her Supreme Champion honors at World Dairy Expo in 1980, where judges declared her “the new ideal Holstein cow.” This breeding masterpiece—out of an EX-91 GMD DOM Pawnee Farm Arlinda Chief dam—later received All-Time All-American Aged Cow recognition in 1984, cementing her place among the greatest show cows in Holstein history. Ella represents the pinnacle of Elevation’s ability to transmit both exceptional type and production traits simultaneously, embodying the “genetic coherence” that made her sire the Bull of the Century. Photo credit: Jack Remsberg.

It wasn’t just Elevation’s sons making history, either. His daughters were equally remarkable. He once led the list for the most Excellent daughters and daughters, scoring 95, 96, and 97 points – the cream of the crop in classification terms. Stars like Ella and Twinkie (both EX-97 All-Time All-Americans), Cora (EX-GMD, dam of Carnation Counselor), and Lindy (EX-GMD, dam of Townson Lindy) didn’t just win in the show ring – they produced sons and grandsons that became influential sires themselves.

Elevation was considered the bull with the most descendants in the United States. It has been found that the two most influential bulls to Holstein US sires were Round Oak Rag Apple Elevation (Elevation) and Pawnee Farm Arlinda Chief (Chief)—up to 99% of AI bulls born after 2010 can be traced back to these animals. There’s something almost poetic about how his genetics have persisted through generation after generation, creating a legacy that continues to shape the Holstein breed today.

Clinton-Camp Majesty (EX-EXTRA), a pivotal son of Round Oak Rag Apple Elevation, photographed in his prime. Though modern genomic metrics undervalue his production (-2,366 lbs milk), Majesty inherited his sire’s structural strengths—including exceptional body strength (+0.97) and sound feet—while passing on key longevity traits to daughters. His genetic profile embodies the Elevation paradox: foundational yet penalized by the same breed progress he enabled. Photo: ST Genetics
Clinton-Camp Majesty (EX-EXTRA), a pivotal son of Round Oak Rag Apple Elevation, photographed in his prime. Though modern genomic metrics undervalue his production, Majesty inherited his sire’s structural strengths—including exceptional body strength and sound feet—while passing on key longevity traits to daughters. His genetic profile embodies the Elevation paradox: foundational yet penalized by the same breed progress he enabled.

A Legacy That Defies Time

You know what’s truly mind-blowing? Six decades after Elevation’s birth, his DNA still runs through 14.5% of active proven Holstein sires. In a world where genetic trends come and go faster than fashion styles, that staying power is nothing short of miraculous.

If you look at Elevation’s current CDCB genomic summary, you might wonder what all the fuss is about. It shows a Net Merit (NM$) of -821 and negative milk production traits (-2,483 lbs milk, -87 lbs fat). But these numbers don’t tell the real story – they’re comparing him to a modern Holstein population he helped create! It’s like penalizing your grandfather for not knowing how to use an iPhone when he helped invent the telephone.

Charlie Will, Holstein Program Manager at Select Sires, speaking at an industry event in his trademark red and black cooperative jacket. With over 45 <a href='https://www.thebullvine.com/a-i-industry/celebrating-50-years-of-semex-a-symbol-of-genetic-progress-and-technological-innovation/' data-lazy-src=

Charlie Will of Select Sires put it perfectly: “Elevation’s genes form the baseline against which we measure progress—you can’t delete the foundation of a skyscraper and expect it to stand.”

What’s fascinating is how specific Elevation traits continue to persist in elite modern genetics:

  • His signature high, wide rear udders (linked to mammary system haplotypes on chromosome 6) remain prevalent in 78% of bulls with >2,000 GTPI.
  • In current evaluations, his descendants inherit body condition scoring alleles associated with +1.1 Livability and +4.5 Daughter Pregnancy Rate.
  • The “Elevation fertility cluster” on chromosome 18 still appears in 63% of high-fertility sires today.

These traits contribute to what breeders call the “Elevation Effect” – cows that maintain production across multiple lactations despite increasing herd turnover rates. His descendants show 18% lower involuntary culling rates than non-elevation lines, making them ideal for pasture-based and robotic milking systems.

Here’s another mind-bender: Elevation’s DNA makes up 8.3% of the CDCB’s genomic reference population. This creates a fascinating paradox where modern genetic evaluations compare new bulls against a baseline that Elevation helped establish. No wonder 80% of elite genomic young sires carry at least one major Elevation haplotype!

Why Elevation Still Matters Today

Today’s breeding programs face a critical choice: preserve Elevation’s durability traits or chase marginal production gains. I’ve talked with farmers who’ve taken the balanced approach, maintaining 12-15% Elevation-derived genetics in their herds. They report some impressive results: 22% lower vet costs, +0.8 lactations per cow, and 3.2% higher lifetime profit than herds chasing the highest genomic numbers.

Elevation’s story paralleled critical advances in reproductive technology, creating a perfect storm of genetic proliferation. His career aligned with breakthroughs in semen freezing and storage that extended viable preservation from days to decades. As AI adoption accelerated worldwide in the 1970s, Elevation’s superior genetics rode this wave of technological diffusion. His career also coincided with the development of computerized progeny testing and record keeping, allowing his impact to be measured more precisely than any bull before him.

If there’s a lesson in Elevation’s story, actual genetic progress isn’t always about extremes – it’s about balance. In an era when genomic selection sometimes emphasizes single traits at the expense of others, Elevation reminds us that the most valuable cattle excel across multiple dimensions. They may not be the most extreme in any category, but they last longer, stay healthier, and ultimately make more money for their owners.

Final Thoughts on a Legend

Elevation’s story isn’t just about genetics – it’s about vision. It’s about two cousins looking beyond immediate results to create something lasting. It’s about recognizing that the most transformative influences sometimes come from unexpected places.

Elevation’s DNA still courses through 14.5% of active Holstein sires six decades later, defying modern genomic evaluations that might dismiss his contribution. While contemporary metrics chase hyperspecialization, his balanced genetic blueprint remains fundamental to functional dairy cattle worldwide.

The contradiction he represents is fascinating: modern genomic models may penalize his alleles for “low” production while simultaneously relying on his chromosome 6 haplotypes as reference points for udder health and efficiency. His descendants continue to excel in diverse systems, showing 18% lower involuntary culling rates and thriving in high-tech robotic facilities and grass-based operations.

As Holstein breeders confront sustainability challenges, Elevation’s legacy offers valuable insights. His balanced genetics align perfectly with modern demands for efficient, lower-carbon dairy systems. Studies show his metabolic efficiency alleles correlate with 4.2% reduced methane output – proving that sometimes old genetics solve new problems!

Ultimately, Round Oak Rag Apple Elevation reminds us that genetic progress isn’t about chasing extremes but finding balance. Elevation didn’t just lift the breed; he gave it wings.

And that, my friend, is the kind of legacy to which we can all raise a glass of milk.

Key Takeaways

  • Elevation’s success demonstrates that transformative genetics often emerge from partnerships that challenge short-term breeding trends, offering lessons for today’s genomic-focused selection methods.
  • His five signature traits formed a “genetic symphony” rarely found in combination—most bulls excel in either production or conformation, while Elevation delivered both without compromise.
  • Beyond individual herds, Elevation reshaped entire breeding organizations. His semen sales helped build Select Sires into a global AI powerhouse, and his genetics standardized Holstein traits across 45 countries.
  • Despite modern genomic evaluations rating him negatively, his chromosome 6 haplotypes remain essential reference points for udder health and efficiency, creating a paradox where his genes form the baseline against which progress is measured.
  • His most enduring legacy may be economic efficiency—herds retaining 12-15% Elevation-derived genetics report 22% lower veterinary costs and longer productive lives than those chasing extreme production traits.

Executive Summary

Round Oak Rag Apple Elevation, born in 1965 on a modest Virginia farm, transformed the dairy industry through his unparalleled genetic transmission of five critical traits: production, udder quality, mobility, fertility, and longevity. Dubbed the “Bull of the Century,” Elevation defied conventional breeding wisdom by emerging from an improbable mating between a questionable sire and a slow-maturing dam. His extraordinary ability to elevate mediocre genetics produced daughters averaging 29,500 pounds of milk (15% above contemporaries) while maintaining exceptional udder structure and extended productive lives. With over 10,000 registered sons and an estimated 8.8 million descendants worldwide, his genetic influence continues six decades later, with his DNA present in 14.5% of active Holstein sires despite the genomics revolution that followed him.

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Round Oak Rag Apple Elevation: The Sire That Took the Dairy Breeding Industry to New Heights – Bullvine Legend Series

Round Oak Rag Apple Elevation

Breeding a legendary dairy sire isn’t automatic. It is not as simple as crossing the right sire with the right dam. However, although it isn’t easy, it does happen.

In one of the most famous cases of all, that of Round Oak Rag Apple Elevation, it is somewhat surprising that the tremendous impact that was to become legendary was not immediately obvious.

It took a little time for the world to recognise his greatness. But, in the end, Round Oak Rag Apple Elevation changed  an entire breed and the dairy industry.

The Breeding of Elevation was Far from a Sure Thing

It would take time for the world to recognize the presence of greatness. Elevation was the product of a great mother, Eve, and a questionable father, Tidy Burke. As it happens, Elevation brought together the best from the Burkes, Rag Apples, Triunes and Winterthur bloodlines.  That is one of the reasons why Elevation would never have been totally missed.  Furthermore, Elevation did many important things well.  There are five that stand out: 1. Production 2. Udders 3. Mobility 4. Fertility and 5. Longevity. Measuring any one or all of these traits shows you what made Elevation special, for these are just a few of the characteristics that contributed to his popularity.  However, Elevation went way beyond mere popularity.  This legendary bull made a tremendous impact on the genetics of the Holstein Breed. He changed dairy organizations.  He affected dairy breeding around the world. The fact that Elevation has 10,000 registered sons in the United States alone speaks volume to both his acceptance and impact.

Was Elevation a Product of Genomics or Ahead of Genomic Times?

Today Elevation would have had hair pulled and submitted to DNA testing.  But in the 1960’s, if you can imagine it, here was a bull entering a sampling program from a slow maturing mother and a never classified father. A father who physically significantly lacked both size and mobility.

George Miller

The mating that resulted in Elevation used the combined knowledge and ‘go for it’ attitude of two men: breeder, Ron Hope from Virginia and his advisor, George Miller.  These two were first cousins, and they started three generations back to produce Elevation.  That is the way it was done in those days.  To arrive at Elevation’s dam, Eve, Hope and Miller stacked three sires: Ivanhoe, Gaiety and General.

By the way, in her early life, Eve carried more condition than normal. This is something that is also seen in Elevation daughters.

It is not any wonder, therefore, that Elevation passed on good fertility, given what we know today about the positive correlation between fertility and body condition score.

After completing a Master of Science degree at Virginia Tech, George Miller spent his career in A.I. starting at the field level, then as a state A.I. manager and eventually as Director of Marketing and Development at Select Sires.  George knew Holsteins, and he had access to bull performance information.  There must have been many interesting discussions between these first cousins about who would be the best mate for Eve, in order to produce a son that could enter A.I.

As mentioned previously, Tidy Burke Elevation, Elevation’s sire, was an ugly duckling but he did produce outstanding daughters.  Four of those daughters earned Honorable Mention All-American Get of Sire.  Today, it is evident that an artful breeder and a top notch A.I. man were indeed able to find the best sire available for Eve.  Remember that these men were making their decisions before the world had ever thought of using DNA information to aid in mating.

Elevation Made an Impact on Organisations

Charlie Will, who is the Holstein Sire Program Manager at Select Sires, gives perspective to the impact that Elevation has had on the company that originally purchased him. “Elevation put Select Sires on the map.  He was so far ahead of all other bulls for his time.  He had exceptional production and amazing type at the same time.” He explains what that meant over time, from the beginning and up to and including the present time. “Elevation made it possible for Select sires to grow as a new company.  Today Elevation still ranks #1 at Holstein USA for the most genes in common among today’s active proven sires (14.5%).  His impact continues 52 years after his birth.”

The WOW Factor of Elevation.

It’s easy to reiterate what set Elevation apart from the competition. Charlie keeps it simple. “Elevation had extraordinary type and production in one package.”  He sees this combination as almost miraculous.  “he dominated the mating no matter what kind of cow you used him on.  He could make a Great Cow from a Poor dam.  This is why he could have a huge impact in a single generation.”

Elevation Didn’t Just INFLUENCE the Future, He MADE the Future!

Breeders always pay attention to cow families.  But in order to influence an entire population, you must go beyond sires and look at their descendants.  Elevation influenced one generation after another: his kids, his kid’s kids, his kid’s kid’s kids.  This is what made Elevation’s influence stellar.

Facts Alone Don’t Spark Legends.  Results Do.

In any business, repeatable results are the only true measure of legendary success.  Popularity and memories fade.  In dairy cattle breeding, generations of descendants tell the real story. 

Charlie Will
Holstein Sire Program Manager
Select Sires

In describing Elevation daughters, Charlie Will starts with a somewhat modest description.  “His daughters had great legs and feet.  A straighter leg but with healthy hocks and strong loins.” Warming to the topic, Charlie adds “Elevation daughters are tall enough, but not extreme, with ideal dairy strength and proportional width for the stature.”  He concludes with what made the difference. “The typical Elevation daughters were short headed heifers but, when they were called into line, their exceptional udders, high and wide Rear Udder, and the great shape and symmetry of their udders, quickly made a breeder proud to own her.”

The first appearance was not always the final answer with Elevation daughters that became long- lived high production cows.

Once proven, everyone recognized that Elevation would continue stamping out great daughters, as he moved the Holstein breed to new heights.

Elevation’s Legacy Lives on Through His Sons and Daughters

Since almost all sires active in the breed today trace back to Elevation, Charlie Will finds it hard to pick from a list where the greats are almost too numerous to mention.  For him, Elevation’s most impact sons include, “Bova, Starbuck, Pete, Mars Tony, Sexation, and Lime Hollow Mars.”

On the daughter side, Charlie lists many attributes. “Elevation has had many class winning daughters, including at World Dairy Expo and the Royal Winter. Elevation also led the list, at one time, for the number of Excellent daughters and also for the number of daughters who scored 95, 96 and 97.” His daughter list reads like an all-star lineup, from EX97 All-Time All-American’s Ella and Twinkie to EX-GMD Cora and Lindy, the dams of Carnation Counselor and Townson Lindy, respectively.

Elevation Surpasses All Heights

When we recognize a dairy legend, it is great to hear some stories from behind the scenes.  Charlie tells one about the time that Elevation was classified 96. “Jim Patterson was head of the Holstein USA classification program at the time that Elevation was raised from 95 to 96.  Later, after he retired, he told me that he only made one mistake, in all the years that he classified. He wished that he would have made Elevation 97 instead of 96!” (Learn more: CHARLIE WILL “A CAREER WITH IMPACT” – SELECT SIRES 50TH ANNIVERSARY)

Northcroft Ella Elevation EX-97-4E
1980 – Grand/Supreme Champion – WDE
1981 – Grand Champion – RAWF
1977, 1980, 1981, 1982 – All-American

Elevation’s Impact is Felt

With the perfect vision accorded to us by hindsight, we can clearly see that Elevation didn’t only influence genetics. Elevation has also had a tremendous impact on sales, new research and the success of countless breeders and organisations. Dairy strategy and development have also felt his influence. And, ultimately, the dairy show ring was also impacted by Elevation.

The World Wide Elevation Influence

Elevation, often known as RORAE, made friends for United States Holsteins around the world.  Therein lies the engine that drives the legend.  Fundamentally, around the world, one bull, through his progeny, significantly changed the profitability of the Holstein cow.  But the measure of Elevation goes beyond mere financial success.  Elevation made many dairymen into successful dairy breeders.  How did he do it? Elevation stamped out daughters that provided what dairymen needed. Production. Longevity. Fertility. Mobility. Functional mammary systems. These are the characteristics passed on by a one-of-a-kind, legendary bull.

The Bullvine Bottom Line

Elevation forever lifted the worldwide dairy breeding industry to a new level.

Greatness can have many definitions, but in Holstein breeding, it can be said using a single name, Round Oak Rag Apple ELEVATION.

 

 

 

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The Genetic Genius of Darwin, Mendel and Hunt – Genetic Transmission and the Holstein Cow

There is no question that when it comes to understanding what cows will transmit and what cows will not, it is an enigma wrapped in a conundrum.  There is much that we don’t know and some would argue it is not meant to be known.  The problem is, for those of us with a passion for breeding great dairy cattle, we want to know it all.  For that I turn to the three greatest genetic geniuses in the history of the world, Darwin, Mendel and Hunt (No they are not a law firm).

Charles Robert Darwin He established that all species of life have descended over time from common ancestors, and proposed the scientific theory that this branching pattern of evolution resulted from a process that he called natural selection, in which the struggle for existence has a similar effect to the artificial selection involved in selective breeding.

Charles Darwin

Ask anyone in the world to name a geneticist and the first name that comes to mind has to be Charles Darwin.  No better demonstration of Darwin’s theory of evolution exists in the world than in dairy cattle breeding.  While there is no question that artificial selection and selective breeding exist on a daily basis, a cow’s ability to reproduce and produce milk leads to a natural level of selection that epitomizes Darwin’s theory.  “The laws governing inheritance,” Darwin wrote, “are for the most part unknown.”  Moreover, while many modern geneticists have theories about the tendencies of the modern Holstein cow, their genetic transmission pathways in large part remain a mystery to this day.

Gregor Mendel

Gregor Mendel

Then along came Gregor Mendel who introduced the concept of “genes” to explain heritability.  Mendel changed the whole way we look at breeding when he introduced the theory that the chromosome is the carrier of genetic traits.  He also explained why a trait can disappear in one generation and reappear in the next and why these traits occur in a three-to-one ratio.  One of Mendel’s disciples, three quarters of a century later, was Thomas B. Macaulay.  Macaulay conducted his own studies, on his Mount Victoria Farms (Read more: Mount Victoria Farms – The art and science of great breeding).

Thomas Hunt Morgan

Thomas Hunt Morgan

Then along came Hunt. Well, more specifically, Thomas Hunt Morgan, but my ego wouldn’t let this go as my name is Andrew Morgan Hunt (Read more about my ego: I’m Sorry But I’ve Had Just About Enough Of… ).  In research that is now reproduced by grade 9 science students around the world, Morgan introduced the concept of X and Y-chromosomes.  Morgan concluded that a female has two X chromosomes and that males have both X and Y-chromosomes.  He also posited that the male of the species, because of the presence of the Y chromosome, transmits differently than the female.

To get a better understanding of this, let’s look at this from both sides of the story.

His side of the story (XY)

If you look at Holstein bulls throughout history you find four distinct patterns:

  1. Great daughters but no legacy sons
    These are the bulls that sired amazing brood cows but none of their sons were able to continue their genetic legacy.  Examples are Hanover-Hill Triple Threat, Carlin-M Ivanhoe Bell, and Braedale Goldwyn.  They all were able to sire brood cow daughters beyond compare, but no real sons to advance that genetic legacy.  Why did these sires seem to produce better on the female side than that of the male?  For that we need to turn to Morgan and his X and Y chromosome theory.  Since the Y chromosome is the only one that is inherited solely via the paternal  line, this leads  some geneticists to believe that it carries little genetic information, and as a result  a great sires genetic legacy rest more with his daughters than with his sons.  Therefore, with this first group of sires it is thought that much of their genetics were transmitted on the X chromosome rather than the Y.
  2. Great sons but not as many brood cows
    Bulls that sired outstanding sons but never produced a top daughter.  A couple of great examples of this are Montvic Rag Apple Sovereign, Maizefield Bellwood and O-Bee Manfred Justice.  All of these sires have left outstanding sons, but are not found as often in the maternal sire stack of the great sires.  There is no question as to their genetic contribution to the breed, but it was more as a sire of sons than their ability to leave an equal number of brood cows.
  3. Sons and daughters both extraordinary
    These are the sires that have gone down in history as the all-time greats.  Sires like Johanna Rag Apple Pabst, Governor of Carnation, Montvic Chieftain, Wisconsin Admiral Burke Lad, A.B.C. Reflection Sovereign, Round Oak Rag Apple Elevation, Pawnee Farm Alrinda Chief, Walkway Chief Mark, Hanoverhill Starbuck, Madawaska Aerostar and Maughlin Storm.  These are the bulls that not only displayed personal greatness but were also able to transmit both outstanding brood cows as well as legacy sons.
  4. Sons and daughters that were inferior
    Sons and daughters that are both below average.  These bulls left inferior daughters and as a result were never even given the chance to produce sons.  Bulls in this category are too numerous to mention and loads of their daughters go to the slaughterhouses every day.  No explanation necessary other than a lack of genetic merit and here enters the need for genomics (Read more: The Truth About Genomic Indexes – “Show Me” That They Work).

Her side the story (XX)

The female side of the story uses the same four distinct groups.

  1. Great daughters but no legacy sons
    These are cows with outstanding female descendants but undistinguished males.  Great examples of these are the cow families of Hanover Hill Papoose, Krull Broker Elegance and Plunshanski Chief Faith.  They all were able to leave outstanding female descendants generation after generation, but were never really able to accomplish the same feat on the male side of the story.
  2. Great sons but not as many brood cows
    These are the cows with potent transmitting sons, but daughters who didn’t outperform the average.  Examples of these are Wylamyna Tidy Kathleen (dam of Sir Bess Tidy and Sir Bess Ormsby Tidy Fobes) Lakefield Fobes Delight (dam of Lakefield Fond Hope, Lakefield Fond Delight Fobes and Carnation Royal Master) and Pawnee Farm Glenvue Beauty (dam of Pawnee Farm Arlinda Chief).  All of these cows had outstanding maternal lines but for some reason were just not able to transmit that legacy through their daughters.
  3. Sons and daughters both extraordinary
    Among the females in this category are Glenridge Citation Roxy, Mil-R-Mor Roxette, Comestar Laurie Sheik, Braedale Gypsy Grand and Snow-N Denises Dellia.
  4. Sons and daughters that were inferior
    Cows who, in terms of influence, failed to produce anything worthwhile.  Blame it on lack of genetics, bad breeding, improper management, or just bad luck, these cows just didn’t influence the breed. We have all seen examples.

The Bullvine Bottom Line

There has never been a clear explanation of why some bloodlines seem to transmit better through maternal lines, others through the paternal, and still others do well in both.  Even genomics does not answer this.  There are high genomic animals that still have these same tendencies.  Maybe if we could genomic test the genes on each chromosome we might find the answers?  Until then Genetic Transmission in the Holstein Cow will remain a mystery.

To read more about this get a copy of The Holstein History by Edward Morwick and read the chapter on Inheritance Patterns.


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