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There is no question that when it comes to understanding what cows will transmit and what cows will not, it is an enigma wrapped in a conundrum.  There is much that we don’t know and some would argue it is not meant to be known.  The problem is, for those of us with a passion for breeding great dairy cattle, we want to know it all.  For that I turn to the three greatest genetic geniuses in the history of the world, Darwin, Mendel and Hunt (No they are not a law firm).

Charles Robert Darwin He established that all species of life have descended over time from common ancestors, and proposed the scientific theory that this branching pattern of evolution resulted from a process that he called natural selection, in which the struggle for existence has a similar effect to the artificial selection involved in selective breeding.

Charles Darwin

Ask anyone in the world to name a geneticist and the first name that comes to mind has to be Charles Darwin.  No better demonstration of Darwin’s theory of evolution exists in the world than in dairy cattle breeding.  While there is no question that artificial selection and selective breeding exist on a daily basis, a cow’s ability to reproduce and produce milk leads to a natural level of selection that epitomizes Darwin’s theory.  “The laws governing inheritance,” Darwin wrote, “are for the most part unknown.”  Moreover, while many modern geneticists have theories about the tendencies of the modern Holstein cow, their genetic transmission pathways in large part remain a mystery to this day.

Gregor Mendel

Gregor Mendel

Then along came Gregor Mendel who introduced the concept of “genes” to explain heritability.  Mendel changed the whole way we look at breeding when he introduced the theory that the chromosome is the carrier of genetic traits.  He also explained why a trait can disappear in one generation and reappear in the next and why these traits occur in a three-to-one ratio.  One of Mendel’s disciples, three quarters of a century later, was Thomas B. Macaulay.  Macaulay conducted his own studies, on his Mount Victoria Farms (Read more: Mount Victoria Farms – The art and science of great breeding).

Thomas Hunt Morgan

Thomas Hunt Morgan

Then along came Hunt. Well, more specifically, Thomas Hunt Morgan, but my ego wouldn’t let this go as my name is Andrew Morgan Hunt (Read more about my ego: I’m Sorry But I’ve Had Just About Enough Of… ).  In research that is now reproduced by grade 9 science students around the world, Morgan introduced the concept of X and Y-chromosomes.  Morgan concluded that a female has two X chromosomes and that males have both X and Y-chromosomes.  He also posited that the male of the species, because of the presence of the Y chromosome, transmits differently than the female.

To get a better understanding of this, let’s look at this from both sides of the story.

His side of the story (XY)

If you look at Holstein bulls throughout history you find four distinct patterns:

  1. Great daughters but no legacy sons
    These are the bulls that sired amazing brood cows but none of their sons were able to continue their genetic legacy.  Examples are Hanover-Hill Triple Threat, Carlin-M Ivanhoe Bell, and Braedale Goldwyn.  They all were able to sire brood cow daughters beyond compare, but no real sons to advance that genetic legacy.  Why did these sires seem to produce better on the female side than that of the male?  For that we need to turn to Morgan and his X and Y chromosome theory.  Since the Y chromosome is the only one that is inherited solely via the paternal  line, this leads  some geneticists to believe that it carries little genetic information, and as a result  a great sires genetic legacy rest more with his daughters than with his sons.  Therefore, with this first group of sires it is thought that much of their genetics were transmitted on the X chromosome rather than the Y.
  2. Great sons but not as many brood cows
    Bulls that sired outstanding sons but never produced a top daughter.  A couple of great examples of this are Montvic Rag Apple Sovereign, Maizefield Bellwood and O-Bee Manfred Justice.  All of these sires have left outstanding sons, but are not found as often in the maternal sire stack of the great sires.  There is no question as to their genetic contribution to the breed, but it was more as a sire of sons than their ability to leave an equal number of brood cows.
  3. Sons and daughters both extraordinary
    These are the sires that have gone down in history as the all-time greats.  Sires like Johanna Rag Apple Pabst, Governor of Carnation, Montvic Chieftain, Wisconsin Admiral Burke Lad, A.B.C. Reflection Sovereign, Round Oak Rag Apple Elevation, Pawnee Farm Alrinda Chief, Walkway Chief Mark, Hanoverhill Starbuck, Madawaska Aerostar and Maughlin Storm.  These are the bulls that not only displayed personal greatness but were also able to transmit both outstanding brood cows as well as legacy sons.
  4. Sons and daughters that were inferior
    Sons and daughters that are both below average.  These bulls left inferior daughters and as a result were never even given the chance to produce sons.  Bulls in this category are too numerous to mention and loads of their daughters go to the slaughterhouses every day.  No explanation necessary other than a lack of genetic merit and here enters the need for genomics (Read more: The Truth About Genomic Indexes – “Show Me” That They Work).

Her side the story (XX)

The female side of the story uses the same four distinct groups.

  1. Great daughters but no legacy sons
    These are cows with outstanding female descendants but undistinguished males.  Great examples of these are the cow families of Hanover Hill Papoose, Krull Broker Elegance and Plunshanski Chief Faith.  They all were able to leave outstanding female descendants generation after generation, but were never really able to accomplish the same feat on the male side of the story.
  2. Great sons but not as many brood cows
    These are the cows with potent transmitting sons, but daughters who didn’t outperform the average.  Examples of these are Wylamyna Tidy Kathleen (dam of Sir Bess Tidy and Sir Bess Ormsby Tidy Fobes) Lakefield Fobes Delight (dam of Lakefield Fond Hope, Lakefield Fond Delight Fobes and Carnation Royal Master) and Pawnee Farm Glenvue Beauty (dam of Pawnee Farm Arlinda Chief).  All of these cows had outstanding maternal lines but for some reason were just not able to transmit that legacy through their daughters.
  3. Sons and daughters both extraordinary
    Among the females in this category are Glenridge Citation Roxy, Mil-R-Mor Roxette, Comestar Laurie Sheik, Braedale Gypsy Grand and Snow-N Denises Dellia.
  4. Sons and daughters that were inferior
    Cows who, in terms of influence, failed to produce anything worthwhile.  Blame it on lack of genetics, bad breeding, improper management, or just bad luck, these cows just didn’t influence the breed. We have all seen examples.

The Bullvine Bottom Line

There has never been a clear explanation of why some bloodlines seem to transmit better through maternal lines, others through the paternal, and still others do well in both.  Even genomics does not answer this.  There are high genomic animals that still have these same tendencies.  Maybe if we could genomic test the genes on each chromosome we might find the answers?  Until then Genetic Transmission in the Holstein Cow will remain a mystery.

To read more about this get a copy of The Holstein History by Edward Morwick and read the chapter on Inheritance Patterns.

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Recently Braedale Goldwyn became the 1st sire ever to produce 1000 Excellent daughters in Canada (Read more – Goldwyn First Ever to 1000 EX in Canada).  At this year’s Royal Winter Fair Goldwyn sired 27% of the animals shown, including Grand, Reserve, Honorable Mention, Intermediate, Reserve Intermediate and Honorable Mention Intermediate Champions (Read More – The 2012 Royal Winter Fair Holstein Show – One of the Greatest Stories Ever Told). It is no surprise that many enthusiasts declare Goldwyn to be the greatest type sire ever.

Grand Champion Selection at the 2012 Royal Winter Fair.  All sired by Braedale Goldwyn

Grand Champion Selection at the 2012 Royal Winter Fair. All sired by Braedale Goldwyn

At the Bullvine we wanted to determine the secret to Goldwyn’s tremendous success.  When you look at Goldwyn’s conformation values for Chest Width (-3) and Body Depth (-4) you don’t instantly assume that he would be such a dominant type sire.  In order to analyze Goldwyn’s type transmitting ability versus other great sires, we looked at his ability to sire the pinnacle of the breed – an Excellent cow.  To make it a level playing field, we looked at sires that had a significant number of their daughters that were 5 years or older.  In other words, that were eligible to go Excellent. Here is what we found.

ERBACRES DAMION23-Jul-001590104962125425774
BRAEDALE GOLDWYN3-Jan-0012851017108371449344502713
SHOREMAR BKB ALLEGRO1-Sep-001188221031363327
WILCOXVIEW JASPER5-Jun-991184159251739261211562
INNWOOD TERRASON21-Jun-96108223815832562914484
LADINO PARK TALENT-IMP31-Mar-9810795174624884335831943
BKB AFFIRMED17-Sep-0110843771444201
RENAISSANCE TRIUMPHANT7-May-9598810144643413159
FUSTEAD EMORY BLITZ2-Mar-969752132118439920621532
SUNNYLODGE LINJET1-Mar-928872823921244365712
EK-OSEEANA ASPEN4-May-00883241011615467
REGANCREST DUNDEE3-May-99883480387157371984914
ROYLANE JORDAN1-May-96881195159125831022474
WINDY-KNOLL-VIEW PRIMETIME8-Mar-007819152946831796
DUPASQUIER CARISMA25-Sep-0178359714137741792
BOSS IRON17-Jul-9677521427948439231
MARKWELL KITE12-Mar-99679776191017432274
VALLEYRIVER RUBEN REDMAN17-Jun-99680918141403536253
INDIANHEAD ENCOUNTER28-Nov-995791720035314872
OSEEANA ASTRONOMICAL15-Sep-9048121911571582638556
HANOVER-HILL-R SPIRIT26-Dec-92280127569783347167
HANOVERHILL STARBUCK26-Mar-79-171419725719324108181771
DONNANDALE SKYCHIEF24-Aug-86-2742772573442924231243
MARSHFIELD ELEVATION TONY6-Aug-72-67336776186996071

What stands out when looking at these sires is that, even though Goldwyn has a high conformation index, there are no less than 12 sires (Linjet, Triumphant, Allegro, Aspen, Spirit, Primetime, Astronomical, Damion, Terrason, Dundee, Jordan and Kite) that actually had a higher percentage of their classified daughters score EX.  Linjet, Triumphant, Allegro, Aspen, Spirit, Primetime, and Astronomical have more than double the percentage of their daughters scoring Excellent compared to Goldwyn.


2012 Supreme Champion World Dairy Expo
2012 Supreme Champion Royal Agriculture Winter Fair

Let’s take a closer look at Linjet, Triumphant, Allegro and Aspen.  These sires not only have a higher percentage of daughters classifying Excellent, they also have a lower percentage scoring Good or lower.  This highlighted something very interesting.  On average, the 2 yr old daughters of this group of sires actually scored lower than Goldwyn’s daughters, but as they matured, they tended to significantly increase in score.  Since the sire conformation index only uses classification scores as 2 year olds, this age improvement was not reflected in all of their conformation scores.  Not being able to foretell this age improvement had a significant effect on semen sales of these bulls because most breeders use conformation to determine what type sires to use.  As a result breeders missed out on these sires whose daughters matured later and scored higher later in life.  This was especially true for Allegro and Aspen, whose later-maturing daughters developed outstanding dairy strength post first lactation.

The Bullvine Bottom Line

There is no question that Goldwyn has had the most number of Excellent daughters, nearly double the number produced by second place sire, LADINO PARK TALENT-IMP.  But why is this? Is it the result of his great type transmitting ability, or is it his type proof combined with his high LPI proof resulted in much greater use.  In other words, is it volume or is it quality?  Undoubtedly,  Goldwyn has been the most dominant type sire of the past decade, the bottom line question is,”Does Goldwyn dominate because of ability or because of opportunity?”

Looking to find out what to breed your Golwyn’s to?  Check this out Breeding the Next Show Winners


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